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==Genetics== |
==Genetics== |
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The use of [[mtDNA|mitochondrial]] "mtDNA" (female lineage) and [[Human Y-chromosome DNA haplogroups|Y-chromosomal]] "Y-DNA" (male lineage) DNA-markers in tracing back the history of human populations has been gaining ground in ethnographic studies of Finnish people (e.g. the National Geographic Genographic Project<ref name="GP">{{cite web|url=https://www3.nationalgeographic.com/genographic/atlas.html|title=Atlas of the Human Journey – The Genographic Project|date=5 April 2008|url-status=unfit |archive-url=https://web.archive.org/web/20080405232839/https://www3.nationalgeographic.com/genographic/atlas.html|archive-date=5 April 2008}}</ref> and the Suomi DNA-projekti.) The most common maternal haplogroup among Finns is [[Haplogroup H (mtDNA)|H]], as 41.5% of Finnish women belong to it. One in four carry the [[Haplogroup U (mtDNA)|haplogroup U5]].<ref name=":3" /> It is estimated to be the oldest major mtDNA haplogroup in Europe and is found in the whole of Europe at a low frequency, but seems to be found in significantly higher levels among Finns, [[Estonians]] and the [[Sami people]].<ref name="GP" /> The older population of European [[hunter-gatherer]]s that lived across large parts of Europe before the early farmers appeared are outside the genetic variation of modern populations, but most similar to Finns.<ref>{{Cite news|url=https://www.aaas.org/news/science-stone-age-skeletons-suggest-europe-s-first-farmers-came-southern-europe|title=Science: Stone Age Skeletons Suggest Europe's First Farmers Came From Southern Europe|date=26 April 2012|work=AAAS – The World's Largest General Scientific Society|access-date=8 August 2017|language=en|archive-date=8 August 2017|archive-url=https://web.archive.org/web/20170808160639/https://www.aaas.org/news/science-stone-age-skeletons-suggest-europe-s-first-farmers-came-southern-europe|url-status=live}}</ref> |
The use of [[mtDNA|mitochondrial]] "mtDNA" (female lineage) and [[Human Y-chromosome DNA haplogroups|Y-chromosomal]] "Y-DNA" (male lineage) DNA-markers in tracing back the history of human populations has been gaining ground in ethnographic studies of Finnish people (e.g. the National Geographic Genographic Project<ref name="GP">{{cite web|url=https://www3.nationalgeographic.com/genographic/atlas.html|title=Atlas of the Human Journey – The Genographic Project|date=5 April 2008|url-status=unfit |archive-url=https://web.archive.org/web/20080405232839/https://www3.nationalgeographic.com/genographic/atlas.html|archive-date=5 April 2008}}</ref> and the Suomi DNA-projekti.) The most common maternal haplogroup among Finns is [[Haplogroup H (mtDNA)|H]], as 41.5% of Finnish women belong to it. One in four carry the [[Haplogroup U (mtDNA)|haplogroup U5]].<ref name=":3" /> It is estimated to be the oldest major mtDNA haplogroup in Europe and is found in the whole of Europe at a low frequency, but seems to be found in significantly higher levels among Finns, [[Estonians]] and the [[Sami people]].<ref name="GP" /> The older population of European [[hunter-gatherer]]s that lived across large parts of Europe before the early farmers appeared are outside the genetic variation of modern populations, but most similar to Finns.<ref>{{Cite news|url=https://www.aaas.org/news/science-stone-age-skeletons-suggest-europe-s-first-farmers-came-southern-europe|title=Science: Stone Age Skeletons Suggest Europe's First Farmers Came From Southern Europe|date=26 April 2012|work=AAAS – The World's Largest General Scientific Society|access-date=8 August 2017|language=en|archive-date=8 August 2017|archive-url=https://web.archive.org/web/20170808160639/https://www.aaas.org/news/science-stone-age-skeletons-suggest-europe-s-first-farmers-came-southern-europe|url-status=live}}</ref> |
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With regard to the Y-chromosome, the most common haplogroups of the Finns is [[Haplogroup N (Y-DNA)#Haplogroup N1c|N1c]], as it is carried by 58–64 percent of Finnish men.<ref name=":11">{{Cite journal |last=Preussner |first=Annina |last2=Leinonen |first2=Jaakko |last3=Riikonen |first3=Juha |last4=Pirinen |first4=Matti |last5=Tukiainen |first5=Taru |date=2024-10-28 |title=Y chromosome sequencing data suggest dual paths of haplogroup N1a1 into Finland |url=https://www.nature.com/articles/s41431-024-01707-7 |journal=European Journal of Human Genetics |language=en |pages=1–9 |doi=10.1038/s41431-024-01707-7 |issn=1476-5438}}</ref> N1c, which is found mainly in a few countries in Europe (Estonia, Finland, Latvia, Lithuania, and Russia), is a subgroup of the [[haplogroup N (Y-DNA)|haplogroup N]] distributed across northern Eurasia and suggested to have entered Europe from Siberia.<ref name="RootsiZhivotovsky2006">{{cite journal|last1=Rootsi|first1=Siiri|last2=Zhivotovsky|first2=Lev A|last3=Baldovič|first3=Marian|last4=Kayser|first4=Manfred|last5=Kutuev|first5=Ildus A|last6=Khusainova|first6=Rita|last7=Bermisheva|first7=Marina A|last8=Gubina|first8=Marina|last9=Fedorova|first9=Sardana A|last10=Ilumäe|first10=Anne-Mai|last11=Khusnutdinova|first11=Elza K|last12=Voevoda|first12=Mikhail I|last13=Osipova|first13=Ludmila P|last14=Stoneking|first14=Mark|last15=Lin|first15=Alice A|last16=Ferak|first16=Vladimir|last17=Parik|first17=Jüri|last18=Kivisild|first18=Toomas|last19=Underhill|first19=Peter A|last20=Villems|first20=Richard|title=A counter-clockwise northern route of the Y-chromosome haplogroup N from Southeast Asia towards Europe|journal=European Journal of Human Genetics|volume=15|issue=2|year=2007|pages=204–211|issn=1018-4813|doi=10.1038/sj.ejhg.5201748|pmid=17149388|doi-access=free}}</ref> The haplogroup N is typical for Uralic-speaking peoples.<ref name=":3" /> Other Y-DNA haplogroups among Finns include [[Haplogroup I (Y-DNA)|I1a]] (25 %), [[Haplogroup R1a (Y-DNA)|R1a]] (4.3 %), and [[Haplogroup R1b (Y-DNA)|R1b]] (3.5 %).<ref name=":11" /> |
With regard to the Y-chromosome, the most common haplogroups of the Finns is [[Haplogroup N (Y-DNA)#Haplogroup N1c|N1c]], as it is carried by 58–64 percent of Finnish men.<ref name=":11">{{Cite journal |last=Preussner |first=Annina |last2=Leinonen |first2=Jaakko |last3=Riikonen |first3=Juha |last4=Pirinen |first4=Matti |last5=Tukiainen |first5=Taru |date=2024-10-28 |title=Y chromosome sequencing data suggest dual paths of haplogroup N1a1 into Finland |url=https://www.nature.com/articles/s41431-024-01707-7 |journal=European Journal of Human Genetics |language=en |pages=1–9 |doi=10.1038/s41431-024-01707-7 |issn=1476-5438}}</ref> N1c, which is found mainly in a few countries in Europe (Estonia, Finland, Latvia, Lithuania, and Russia), is a subgroup of the [[haplogroup N (Y-DNA)|haplogroup N]] distributed across northern Eurasia and suggested to have entered Europe from Siberia.<ref name="RootsiZhivotovsky2006">{{cite journal|last1=Rootsi|first1=Siiri|last2=Zhivotovsky|first2=Lev A|last3=Baldovič|first3=Marian|last4=Kayser|first4=Manfred|last5=Kutuev|first5=Ildus A|last6=Khusainova|first6=Rita|last7=Bermisheva|first7=Marina A|last8=Gubina|first8=Marina|last9=Fedorova|first9=Sardana A|last10=Ilumäe|first10=Anne-Mai|last11=Khusnutdinova|first11=Elza K|last12=Voevoda|first12=Mikhail I|last13=Osipova|first13=Ludmila P|last14=Stoneking|first14=Mark|last15=Lin|first15=Alice A|last16=Ferak|first16=Vladimir|last17=Parik|first17=Jüri|last18=Kivisild|first18=Toomas|last19=Underhill|first19=Peter A|last20=Villems|first20=Richard|title=A counter-clockwise northern route of the Y-chromosome haplogroup N from Southeast Asia towards Europe|journal=European Journal of Human Genetics|volume=15|issue=2|year=2007|pages=204–211|issn=1018-4813|doi=10.1038/sj.ejhg.5201748|pmid=17149388|doi-access=free}}</ref> The haplogroup N is typical for Uralic-speaking peoples.<ref name=":3" /><ref name=":10" /> Other Y-DNA haplogroups among Finns include [[Haplogroup I (Y-DNA)|I1a]] (25 %), [[Haplogroup R1a (Y-DNA)|R1a]] (4.3 %), and [[Haplogroup R1b (Y-DNA)|R1b]] (3.5 %).<ref name=":11" /> |
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Finns are genetically closest to [[Karelians]], a fellow Balto-Finnic group.<ref>Lang, Valter (2020). ''Homo Fennicus – Itämerensuomalaisten etnohistoria''. Finnish Literature Society. p. 95. {{ISBN|978-951-858-130-0}}</ref><ref name=":3" /> Finns and Karelians form a cluster with another Balto-Finnic people, the [[Vepsians|Veps]].<ref name=":2">{{Cite journal |last1=Zhernakova |first1=Daria V. |last2=Brukhin |first2=Vladimir |last3=Malov |first3=Sergey |last4=Oleksyk |first4=Taras K. |last5=Koepfli |first5=Klaus Peter |last6=Zhuk |first6=Anna |last7=Dobrynin |first7=Pavel |last8=Kliver |first8=Sergei |last9=Cherkasov |first9=Nikolay |last10=Tamazian |first10=Gaik |last11=Rotkevich |first11=Mikhail |last12=Krasheninnikova |first12=Ksenia |last13=Evsyukov |first13=Igor |last14=Sidorov |first14=Sviatoslav |last15=Gorbunova |first15=Anna |date=January 2020 |title=Genome-wide sequence analyses of ethnic populations across Russia |journal=Genomics |volume=112 |issue=1 |pages=442–458 |doi=10.1016/j.ygeno.2019.03.007 |issn=1089-8646 |pmid=30902755 |s2cid=85455300|url=https://pure.rug.nl/ws/files/118806867/1_s2.0_S0888754318307419_main.pdf }}</ref><ref name=":3" /><ref>{{Cite journal |last1=Pankratov |first1=Vasili |last2=Montinaro |first2=Francesco |last3=Kushniarevich |first3=Alena |last4=Hudjashov |first4=Georgi |last5=Jay |first5=Flora |last6=Saag |first6=Lauri |last7=Flores |first7=Rodrigo |last8=Marnetto |first8=Davide |last9=Seppel |first9=Marten |last10=Kals |first10=Mart |last11=Võsa |first11=Urmo |last12=Taccioli |first12=Cristian |last13=Möls |first13=Märt |last14=Milani |first14=Lili |last15=Aasa |first15=Anto |date=25 July 2020 |title=Differences in local population history at the finest level: the case of the Estonian population |journal=European Journal of Human Genetics |language=en |volume=28 |issue=11 |pages=1580–1591 |doi=10.1038/s41431-020-0699-4 |pmid=32712624 |pmc=7575549 |issn=1476-5438}}</ref> They show relative affinity to Northern Russians as well,<ref name=":9">{{Cite journal |last1=Kidd |first1=Kenneth K. |last2=Evsanaa |first2=Baigalmaa |last3=Togtokh |first3=Ariunaa |last4=Brissenden |first4=Jane E. |last5=Roscoe |first5=Janet M. |last6=Dogan |first6=Mustafa |last7=Neophytou |first7=Pavlos I. |last8=Gurkan |first8=Cemal |last9=Bulbul |first9=Ozlem |last10=Cherni |first10=Lotfi |last11=Speed |first11=William C. |last12=Murtha |first12=Michael |last13=Kidd |first13=Judith R. |last14=Pakstis |first14=Andrew J. |date=4 May 2022 |title=North Asian population relationships in a global context |journal=Scientific Reports |language=en |volume=12 |issue=1 |page=7214 |doi=10.1038/s41598-022-10706-x |issn=2045-2322 |pmc=9068624 |pmid=35508562|bibcode=2022NatSR..12.7214K }}</ref><ref name=":2" /><ref>{{Cite journal |last1=Kushniarevich |first1=Alena |last2=Utevska |first2=Olga |last3=Chuhryaeva |first3=Marina |last4=Agdzhoyan |first4=Anastasia |last5=Dibirova |first5=Khadizhat |last6=Uktveryte |first6=Ingrida |last7=Möls |first7=Märt |last8=Mulahasanovic |first8=Lejla |last9=Pshenichnov |first9=Andrey |last10=Frolova |first10=Svetlana |last11=Shanko |first11=Andrey |last12=Metspalu |first12=Ene |last13=Reidla |first13=Maere |last14=Tambets |first14=Kristiina |last15=Tamm |first15=Erika |date=2 September 2015 |title=Genetic Heritage of the Balto-Slavic Speaking Populations: A Synthesis of Autosomal, Mitochondrial and Y-Chromosomal Data |journal=PLOS ONE |language=en |volume=10 |issue=9 |pages=e0135820 |doi=10.1371/journal.pone.0135820 |pmid=26332464 |pmc=4558026 |bibcode=2015PLoSO..1035820K |issn=1932-6203 |doi-access=free }}</ref> who are known to be at least partially descended from Finno-Ugric-speakers who inhabited Northwestern Russia before the Slavs.<ref name=":3" /><ref>{{Cite journal |last1=Peltola |first1=Sanni |last2=Majander |first2=Kerttu |last3=Makarov |first3=Nikolaj |last4=Dobrovolskaya |first4=Maria |last5=Nordqvist |first5=Kerkko |last6=Salmela |first6=Elina |last7=Onkamo |first7=Päivi |date=9 January 2023 |title=Genetic admixture and language shift in the medieval Volga-Oka interfluve |journal=Current Biology |volume=33 |issue=1 |pages=174–182.e10 |doi=10.1016/j.cub.2022.11.036 |issn=0960-9822|doi-access=free |pmid=36513080 }}</ref> |
Finns are genetically closest to [[Karelians]], a fellow Balto-Finnic group.<ref>Lang, Valter (2020). ''Homo Fennicus – Itämerensuomalaisten etnohistoria''. Finnish Literature Society. p. 95. {{ISBN|978-951-858-130-0}}</ref><ref name=":3" /> Finns and Karelians form a cluster with another Balto-Finnic people, the [[Vepsians|Veps]].<ref name=":2">{{Cite journal |last1=Zhernakova |first1=Daria V. |last2=Brukhin |first2=Vladimir |last3=Malov |first3=Sergey |last4=Oleksyk |first4=Taras K. |last5=Koepfli |first5=Klaus Peter |last6=Zhuk |first6=Anna |last7=Dobrynin |first7=Pavel |last8=Kliver |first8=Sergei |last9=Cherkasov |first9=Nikolay |last10=Tamazian |first10=Gaik |last11=Rotkevich |first11=Mikhail |last12=Krasheninnikova |first12=Ksenia |last13=Evsyukov |first13=Igor |last14=Sidorov |first14=Sviatoslav |last15=Gorbunova |first15=Anna |date=January 2020 |title=Genome-wide sequence analyses of ethnic populations across Russia |journal=Genomics |volume=112 |issue=1 |pages=442–458 |doi=10.1016/j.ygeno.2019.03.007 |issn=1089-8646 |pmid=30902755 |s2cid=85455300|url=https://pure.rug.nl/ws/files/118806867/1_s2.0_S0888754318307419_main.pdf }}</ref><ref name=":3" /><ref>{{Cite journal |last1=Pankratov |first1=Vasili |last2=Montinaro |first2=Francesco |last3=Kushniarevich |first3=Alena |last4=Hudjashov |first4=Georgi |last5=Jay |first5=Flora |last6=Saag |first6=Lauri |last7=Flores |first7=Rodrigo |last8=Marnetto |first8=Davide |last9=Seppel |first9=Marten |last10=Kals |first10=Mart |last11=Võsa |first11=Urmo |last12=Taccioli |first12=Cristian |last13=Möls |first13=Märt |last14=Milani |first14=Lili |last15=Aasa |first15=Anto |date=25 July 2020 |title=Differences in local population history at the finest level: the case of the Estonian population |journal=European Journal of Human Genetics |language=en |volume=28 |issue=11 |pages=1580–1591 |doi=10.1038/s41431-020-0699-4 |pmid=32712624 |pmc=7575549 |issn=1476-5438}}</ref> They show relative affinity to Northern Russians as well,<ref name=":9">{{Cite journal |last1=Kidd |first1=Kenneth K. |last2=Evsanaa |first2=Baigalmaa |last3=Togtokh |first3=Ariunaa |last4=Brissenden |first4=Jane E. |last5=Roscoe |first5=Janet M. |last6=Dogan |first6=Mustafa |last7=Neophytou |first7=Pavlos I. |last8=Gurkan |first8=Cemal |last9=Bulbul |first9=Ozlem |last10=Cherni |first10=Lotfi |last11=Speed |first11=William C. |last12=Murtha |first12=Michael |last13=Kidd |first13=Judith R. |last14=Pakstis |first14=Andrew J. |date=4 May 2022 |title=North Asian population relationships in a global context |journal=Scientific Reports |language=en |volume=12 |issue=1 |page=7214 |doi=10.1038/s41598-022-10706-x |issn=2045-2322 |pmc=9068624 |pmid=35508562|bibcode=2022NatSR..12.7214K }}</ref><ref name=":2" /><ref>{{Cite journal |last1=Kushniarevich |first1=Alena |last2=Utevska |first2=Olga |last3=Chuhryaeva |first3=Marina |last4=Agdzhoyan |first4=Anastasia |last5=Dibirova |first5=Khadizhat |last6=Uktveryte |first6=Ingrida |last7=Möls |first7=Märt |last8=Mulahasanovic |first8=Lejla |last9=Pshenichnov |first9=Andrey |last10=Frolova |first10=Svetlana |last11=Shanko |first11=Andrey |last12=Metspalu |first12=Ene |last13=Reidla |first13=Maere |last14=Tambets |first14=Kristiina |last15=Tamm |first15=Erika |date=2 September 2015 |title=Genetic Heritage of the Balto-Slavic Speaking Populations: A Synthesis of Autosomal, Mitochondrial and Y-Chromosomal Data |journal=PLOS ONE |language=en |volume=10 |issue=9 |pages=e0135820 |doi=10.1371/journal.pone.0135820 |pmid=26332464 |pmc=4558026 |bibcode=2015PLoSO..1035820K |issn=1932-6203 |doi-access=free }}</ref> who are known to be at least partially descended from Finno-Ugric-speakers who inhabited Northwestern Russia before the Slavs.<ref name=":3" /><ref>{{Cite journal |last1=Peltola |first1=Sanni |last2=Majander |first2=Kerttu |last3=Makarov |first3=Nikolaj |last4=Dobrovolskaya |first4=Maria |last5=Nordqvist |first5=Kerkko |last6=Salmela |first6=Elina |last7=Onkamo |first7=Päivi |date=9 January 2023 |title=Genetic admixture and language shift in the medieval Volga-Oka interfluve |journal=Current Biology |volume=33 |issue=1 |pages=174–182.e10 |doi=10.1016/j.cub.2022.11.036 |issn=0960-9822|doi-access=free |pmid=36513080 }}</ref> |
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⚫ | [[File:PCA plot of European individuals.png|thumb|PCA plot of several European populations.<ref name=":4">{{Cite journal |last1=Nelis |first1=Mari |last2=Esko |first2=Tõnu |last3=Mägi |first3=Reedik |last4=Zimprich |first4=Fritz |last5=Zimprich |first5=Alexander |last6=Toncheva |first6=Draga |last7=Karachanak |first7=Sena |last8=Piskáčková |first8=Tereza |last9=Balaščák |first9=Ivan |last10=Peltonen |first10=Leena |last11=Jakkula |first11=Eveliina |date=8 May 2009 |title=Genetic Structure of Europeans: A View from the North–East |journal=PLOS ONE |language=en |volume=4 |issue=5 |pages=e5472 |bibcode=2009PLoSO...4.5472N |doi=10.1371/journal.pone.0005472 |issn=1932-6203 |pmc=2675054 |pmid=19424496 |doi-access=free}}</ref>]] |
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When not compared to these groups, Finns have been found to cluster apart from their neighboring populations, forming outlier clusters.<ref name=":5">{{cite journal |last1=Översti |first1=Sanni |last2=Majander |first2=Kerttu |last3=Salmela |first3=Elina |last4=Salo |first4=Kati |last5=Arppe |first5=Laura |last6=Belskiy |first6=Stanislav |last7=Etu-Sihvola |first7=Heli |last8=Laakso |first8=Ville |last9=Mikkola |first9=Esa |last10=Pfrengle |first10=Saskia |last11=Putkonen |first11=Mikko |last12=Taavitsainen |first12=Jussi-Pekka |last13=Vuoristo |first13=Katja |last14=Wessman |first14=Anna |last15=Sajantila |first15=Antti |date=15 November 2019 |title=Översti, S., Majander, K., Salmela, E. et al. Human mitochondrial DNA lineages in Iron-Age Fennoscandia suggest incipient admixture and eastern introduction of farming-related maternal ancestry. Sci Rep 9, 16883 (2019). |journal=Scientific Reports |volume=9 |issue=1 |page=16883 |doi=10.1038/s41598-019-51045-8 |pmc=6858343 |pmid=31729399 |quote=Finns are also genetically distinct from their neighboring populations and form outliers in the genetic variation within Europe. This genetic uniqueness derives from both reduced genetic diversity and an Asian influence to the gene pool. |last16=Oinonen |first16=Markku |last17=Haak |first17=Wolfgang |last18=Schuenemann |first18=Verena J. |last19=Krause |first19=Johannes |last20=Palo |first20=Jukka U. |last21=Onkamo |first21=Päivi}}</ref> They are shifted away from the cline that most Europeans belong to<ref>{{Cite journal |last1=Lao |first1=Oscar |last2=Lu |first2=Timothy T. |last3=Nothnagel |first3=Michael |last4=Junge |first4=Olaf |last5=Freitag-Wolf |first5=Sandra |last6=Caliebe |first6=Amke |last7=Balascakova |first7=Miroslava |last8=Bertranpetit |first8=Jaume |last9=Bindoff |first9=Laurence A. |last10=Comas |first10=David |last11=Holmlund |first11=Gunilla |last12=Kouvatsi |first12=Anastasia |last13=Macek |first13=Milan |last14=Mollet |first14=Isabelle |last15=Parson |first15=Walther |date=2008 |title=Correlation between Genetic and Geographic Structure in Europe |journal=Current Biology |volume=18 |issue=16 |pages=1241–1248 |doi=10.1016/j.cub.2008.07.049 |pmid=18691889 |s2cid=16945780 |issn=0960-9822|doi-access=free }}</ref> towards geographically distant Uralic-speakers like the [[Udmurts]] and [[Mari people|Mari]] (while remaining genetically distant from them as well).<ref>{{Cite journal |last1=Santos |first1=Patrícia |last2=Gonzàlez-Fortes |first2=Gloria |last3=Trucchi |first3=Emiliano |last4=Ceolin |first4=Andrea |last5=Cordoni |first5=Guido |last6=Guardiano |first6=Cristina |last7=Longobardi |first7=Giuseppe |last8=Barbujani |first8=Guido |date=2020 |title=More Rule than Exception: Parallel Evidence of Ancient Migrations in Grammars and Genomes of Finno-Ugric Speakers |journal=Genes |language=en |volume=11 |issue=12 |page=1491 |doi=10.3390/genes11121491 |pmid=33322364 |pmc=7763979 |issn=2073-4425 |doi-access=free }}</ref> Their distance from Western Europeans is about the same as their distance from [[Komi peoples|Komis]].<ref name=":9" /> The Balto-Finnic Estonians are among the genetically closest populations of Finns, but they are less isolated from the European cluster than Finns. [[Swedes]], while being distinct from the Finns, are also closer to Finns than most European populations.<ref name=":4"/><ref name="Salmela2011">{{cite journal |author=Salmela |display-authors=etal |year=2011 |title=Swedish Population Substructure Revealed by Genome-Wide Single Nucleotide Polymorphism Data |journal=PLOS ONE |volume=6 |issue=2 |pages=e16747 |bibcode=2011PLoSO...616747S |doi=10.1371/journal.pone.0016747 |pmc=3036708 |pmid=21347369 |doi-access=free}}</ref><ref name="Jakkula2008">{{cite journal |author=Jakkula |display-authors=etal |year=2008 |title=The Genome-wide Patterns of Variation Expose Significant Substructure in a Founder Population |journal=The American Journal of Human Genetics |volume=83 |issue=6 |pages=787–794 |doi=10.1016/j.ajhg.2008.11.005 |pmc=2668058 |pmid=19061986}}</ref> |
When not compared to these groups, Finns have been found to cluster apart from their neighboring populations, forming outlier clusters.<ref name=":5">{{cite journal |last1=Översti |first1=Sanni |last2=Majander |first2=Kerttu |last3=Salmela |first3=Elina |last4=Salo |first4=Kati |last5=Arppe |first5=Laura |last6=Belskiy |first6=Stanislav |last7=Etu-Sihvola |first7=Heli |last8=Laakso |first8=Ville |last9=Mikkola |first9=Esa |last10=Pfrengle |first10=Saskia |last11=Putkonen |first11=Mikko |last12=Taavitsainen |first12=Jussi-Pekka |last13=Vuoristo |first13=Katja |last14=Wessman |first14=Anna |last15=Sajantila |first15=Antti |date=15 November 2019 |title=Översti, S., Majander, K., Salmela, E. et al. Human mitochondrial DNA lineages in Iron-Age Fennoscandia suggest incipient admixture and eastern introduction of farming-related maternal ancestry. Sci Rep 9, 16883 (2019). |journal=Scientific Reports |volume=9 |issue=1 |page=16883 |doi=10.1038/s41598-019-51045-8 |pmc=6858343 |pmid=31729399 |quote=Finns are also genetically distinct from their neighboring populations and form outliers in the genetic variation within Europe. This genetic uniqueness derives from both reduced genetic diversity and an Asian influence to the gene pool. |last16=Oinonen |first16=Markku |last17=Haak |first17=Wolfgang |last18=Schuenemann |first18=Verena J. |last19=Krause |first19=Johannes |last20=Palo |first20=Jukka U. |last21=Onkamo |first21=Päivi}}</ref> They are shifted away from the cline that most Europeans belong to<ref>{{Cite journal |last1=Lao |first1=Oscar |last2=Lu |first2=Timothy T. |last3=Nothnagel |first3=Michael |last4=Junge |first4=Olaf |last5=Freitag-Wolf |first5=Sandra |last6=Caliebe |first6=Amke |last7=Balascakova |first7=Miroslava |last8=Bertranpetit |first8=Jaume |last9=Bindoff |first9=Laurence A. |last10=Comas |first10=David |last11=Holmlund |first11=Gunilla |last12=Kouvatsi |first12=Anastasia |last13=Macek |first13=Milan |last14=Mollet |first14=Isabelle |last15=Parson |first15=Walther |date=2008 |title=Correlation between Genetic and Geographic Structure in Europe |journal=Current Biology |volume=18 |issue=16 |pages=1241–1248 |doi=10.1016/j.cub.2008.07.049 |pmid=18691889 |s2cid=16945780 |issn=0960-9822|doi-access=free }}</ref> towards geographically distant Uralic-speakers like the [[Udmurts]] and [[Mari people|Mari]] (while remaining genetically distant from them as well).<ref>{{Cite journal |last1=Santos |first1=Patrícia |last2=Gonzàlez-Fortes |first2=Gloria |last3=Trucchi |first3=Emiliano |last4=Ceolin |first4=Andrea |last5=Cordoni |first5=Guido |last6=Guardiano |first6=Cristina |last7=Longobardi |first7=Giuseppe |last8=Barbujani |first8=Guido |date=2020 |title=More Rule than Exception: Parallel Evidence of Ancient Migrations in Grammars and Genomes of Finno-Ugric Speakers |journal=Genes |language=en |volume=11 |issue=12 |page=1491 |doi=10.3390/genes11121491 |pmid=33322364 |pmc=7763979 |issn=2073-4425 |doi-access=free }}</ref> Their distance from Western Europeans is about the same as their distance from [[Komi peoples|Komis]].<ref name=":9" /> The Balto-Finnic Estonians are among the genetically closest populations of Finns, but they are less isolated from the European cluster than Finns. [[Swedes]], while being distinct from the Finns, are also closer to Finns than most European populations.<ref name=":4"/><ref name="Salmela2011">{{cite journal |author=Salmela |display-authors=etal |year=2011 |title=Swedish Population Substructure Revealed by Genome-Wide Single Nucleotide Polymorphism Data |journal=PLOS ONE |volume=6 |issue=2 |pages=e16747 |bibcode=2011PLoSO...616747S |doi=10.1371/journal.pone.0016747 |pmc=3036708 |pmid=21347369 |doi-access=free}}</ref><ref name="Jakkula2008">{{cite journal |author=Jakkula |display-authors=etal |year=2008 |title=The Genome-wide Patterns of Variation Expose Significant Substructure in a Founder Population |journal=The American Journal of Human Genetics |volume=83 |issue=6 |pages=787–794 |doi=10.1016/j.ajhg.2008.11.005 |pmc=2668058 |pmid=19061986}}</ref> |
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⚫ | Finns being an outlier population has to do with their gene pool having reduced diversity<ref name=":5" /> and differences in admixture, including Asian influence, compared to most Europeans.<ref name=":5" /><ref name=":10" /> In general, Europeans can be modelled to have three ancestral components (hunter-gatherer, farmer and steppe), but this model does not work as such for some northeastern European populations, like the Finns and the Sami.<ref name=":7" /> While their genome is still mostly European, they also have some additional East Eurasian ancestry (varies from 5 up to 10<ref>{{Cite web |last=Nousiainen |first=Anu |date=2018 |title=Keitä me olemme? |url=https://dynamic.hs.fi/2018/dna/ |url-status=live |archive-url=https://web.archive.org/web/20221014035130/https://dynamic.hs.fi/2018/dna/ |archive-date=14 October 2022 |access-date=1 August 2023 |website=HS.fi |language=fi}}</ref>–13<ref>{{Cite journal |last1=Qin |first1=Pengfei |last2=Zhou |first2=Ying |last3=Lou |first3=Haiyi |last4=Lu |first4=Dongsheng |last5=Yang |first5=Xiong |last6=Wang |first6=Yuchen |last7=Jin |first7=Li |last8=Chung |first8=Yeun-Jun |last9=Xu |first9=Shuhua |date=2 April 2015 |title=Quantitating and Dating Recent Gene Flow between European and East Asian Populations |journal=Scientific Reports |language=en |volume=5 |issue=1 |page=9500 |doi=10.1038/srep09500 |issn=2045-2322 |pmc=4382708 |pmid=25833680|bibcode=2015NatSR...5E9500Q }}</ref> % in Finns). This component is most likely [[Siberia|Siberian]]-related, best represented by the north Siberian [[Nganasan people|Nganasans]]. The specific Siberian-like ancestry is suggested to have arrived in Northern Europe during the early [[Iron Age]], linked to the arrival of [[Uralic languages]].<ref name=":3">{{Cite journal |last1=Tambets |first1=Kristiina |last2=Yunusbayev |first2=Bayazit |last3=Hudjashov |first3=Georgi |last4=Ilumäe |first4=Anne-Mai |last5=Rootsi |first5=Siiri |last6=Honkola |first6=Terhi |last7=Vesakoski |first7=Outi |last8=Atkinson |first8=Quentin |last9=Skoglund |first9=Pontus |last10=Kushniarevich |first10=Alena |last11=Litvinov |first11=Sergey |date=21 September 2018 |title=Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations |journal=Genome Biology |volume=19 |issue=1 |page=139 |doi=10.1186/s13059-018-1522-1 |issn=1474-7596 |pmc=6151024 |pmid=30241495 |quote= |doi-access=free }}</ref><ref name=":7">{{Cite journal|last1=Lamnidis|first1=Thiseas C.|last2=Majander|first2=Kerttu|last3=Jeong|first3=Choongwon|last4=Salmela|first4=Elina|last5=Wessman|first5=Anna|last6=Moiseyev|first6=Vyacheslav|last7=Khartanovich|first7=Valery|last8=Balanovsky|first8=Oleg|last9=Ongyerth|first9=Matthias|last10=Weihmann|first10=Antje|last11=Sajantila|first11=Antti|date=27 November 2018|title=Ancient Fennoscandian genomes reveal origin and spread of Siberian ancestry in Europe|journal=Nature Communications|language=en|volume=9|issue=1|page=5018|bibcode=2018NatCo...9.5018L|doi=10.1038/s41467-018-07483-5|issn=2041-1723|pmc=6258758|pmid=30479341|quote=This model, however, does not fit well for present-day populations from north-eastern Europe such as Saami, Russians, Mordovians, Chuvash, Estonians, Hungarians, and Finns: they carry additional ancestry seen as increased allele sharing with modern East Asian populations1,3,9,10. Additionally, within the Bolshoy population, we observe the derived allele of rs3827760 in the EDAR gene, which is found in near-fixation in East Asian and Native American populations today, but is extremely rare elsewhere37, and has been linked to phenotypes related to tooth shape38 and hair morphology39 (Supplementary Data 2). To further test differential relatedness with Nganasan in European populations and in the ancient individuals in this study, we calculated f4(Mbuti, Nganasan; Lithuanian, Test) (Fig. 3). Consistent with f3-statistics above, all the ancient individuals and modern Finns, Saami, Mordovians and Russians show excess allele sharing with Nganasan when used as Test populations.}}</ref> Finns have high steppe or [[Corded Ware culture]]-like admixture, and they have less farmer-related and more hunter-gatherer-related admixture than Scandinavians and West and Central Europeans.<ref name=":10">{{Cite journal |last=Salmela |first=Elina |date=2023 |title=Mistä suomalaisten perimä on peräisin? |url=https://researchportal.helsinki.fi/en/publications/mist%C3%A4-suomalaisten-perim%C3%A4-on-per%C3%A4isin |journal=Duodecim |volume=139 |issue=16 |pages=1247–1255 |issn=0012-7183}}</ref>[[File:Share of 1–2 cM IBD segments of Uralic speakers.png|left|thumb|Share of 1–2 cM IBD segments of Uralic speakers.<ref name=":3" />]]Finns share more [[Identity by descent|identity-by-descent]] (IBD) segments with several other Uralic-speaking peoples, including groups like Estonians, the Sami and the geographically distant Komis and Nganasans, than with their Indo-European-speaking neighbours. This is consistent with the idea that the Uralic peoples share common roots to some degree.<ref name=":3" /> |
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[[File:Share of 1–2 cM IBD segments of Uralic speakers.png|left|thumb|Share of 1–2 cM IBD segments of Uralic speakers (Tambets et al. 2018).]] |
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⚫ | Finns being an outlier population has to do with their gene pool having reduced diversity<ref name=":5" /> and differences in admixture, including Asian influence, compared to most Europeans.<ref name=":5" /><ref name=":10" /> In general, Europeans can be modelled to have three ancestral components (hunter-gatherer, farmer and steppe), but this model does not work as such for some northeastern European populations, like the Finns and the Sami.<ref name=":7" /> While their genome is still mostly European, they also have some additional East Eurasian ancestry (varies from 5 up to 10<ref>{{Cite web |last=Nousiainen |first=Anu |date=2018 |title=Keitä me olemme? |url=https://dynamic.hs.fi/2018/dna/ |url-status=live |archive-url=https://web.archive.org/web/20221014035130/https://dynamic.hs.fi/2018/dna/ |archive-date=14 October 2022 |access-date=1 August 2023 |website=HS.fi |language=fi}}</ref>–13<ref>{{Cite journal |last1=Qin |first1=Pengfei |last2=Zhou |first2=Ying |last3=Lou |first3=Haiyi |last4=Lu |first4=Dongsheng |last5=Yang |first5=Xiong |last6=Wang |first6=Yuchen |last7=Jin |first7=Li |last8=Chung |first8=Yeun-Jun |last9=Xu |first9=Shuhua |date=2 April 2015 |title=Quantitating and Dating Recent Gene Flow between European and East Asian Populations |journal=Scientific Reports |language=en |volume=5 |issue=1 |page=9500 |doi=10.1038/srep09500 |issn=2045-2322 |pmc=4382708 |pmid=25833680|bibcode=2015NatSR...5E9500Q }}</ref> % in Finns). This component is most likely [[Siberia|Siberian]]-related, best represented by the north Siberian [[Nganasan people|Nganasans]]. The specific Siberian-like ancestry is suggested to have arrived in Northern Europe during the early [[Iron Age]], linked to the arrival of [[Uralic languages]].<ref name=":3">{{Cite journal |last1=Tambets |first1=Kristiina |last2=Yunusbayev |first2=Bayazit |last3=Hudjashov |first3=Georgi |last4=Ilumäe |first4=Anne-Mai |last5=Rootsi |first5=Siiri |last6=Honkola |first6=Terhi |last7=Vesakoski |first7=Outi |last8=Atkinson |first8=Quentin |last9=Skoglund |first9=Pontus |last10=Kushniarevich |first10=Alena |last11=Litvinov |first11=Sergey |date=21 September 2018 |title=Genes reveal traces of common recent demographic history for most of the Uralic-speaking populations |journal=Genome Biology |volume=19 |issue=1 |page=139 |doi=10.1186/s13059-018-1522-1 |issn=1474-7596 |pmc=6151024 |pmid=30241495 |quote= |doi-access=free }}</ref><ref name=":7">{{Cite journal|last1=Lamnidis|first1=Thiseas C.|last2=Majander|first2=Kerttu|last3=Jeong|first3=Choongwon|last4=Salmela|first4=Elina|last5=Wessman|first5=Anna|last6=Moiseyev|first6=Vyacheslav|last7=Khartanovich|first7=Valery|last8=Balanovsky|first8=Oleg|last9=Ongyerth|first9=Matthias|last10=Weihmann|first10=Antje|last11=Sajantila|first11=Antti|date=27 November 2018|title=Ancient Fennoscandian genomes reveal origin and spread of Siberian ancestry in Europe|journal=Nature Communications|language=en|volume=9|issue=1|page=5018|bibcode=2018NatCo...9.5018L|doi=10.1038/s41467-018-07483-5|issn=2041-1723|pmc=6258758|pmid=30479341|quote=This model, however, does not fit well for present-day populations from north-eastern Europe such as Saami, Russians, Mordovians, Chuvash, Estonians, Hungarians, and Finns: they carry additional ancestry seen as increased allele sharing with modern East Asian populations1,3,9,10. Additionally, within the Bolshoy population, we observe the derived allele of rs3827760 in the EDAR gene, which is found in near-fixation in East Asian and Native American populations today, but is extremely rare elsewhere37, and has been linked to phenotypes related to tooth shape38 and hair morphology39 (Supplementary Data 2). To further test differential relatedness with Nganasan in European populations and in the ancient individuals in this study, we calculated f4(Mbuti, Nganasan; Lithuanian, Test) (Fig. 3). Consistent with f3-statistics above, all the ancient individuals and modern Finns, Saami, Mordovians and Russians show excess allele sharing with Nganasan when used as Test populations.}}</ref> Finns have high steppe or [[Corded Ware culture]]-like admixture, and they have less farmer-related and more hunter-gatherer-related admixture than Scandinavians and West and Central Europeans.<ref name=":10">{{Cite journal |last=Salmela |first=Elina |date=2023 |title=Mistä suomalaisten perimä on peräisin? |url=https://researchportal.helsinki.fi/en/publications/mist%C3%A4-suomalaisten-perim%C3%A4-on-per%C3%A4isin |journal=Duodecim |volume=139 |issue=16 |pages=1247–1255 |issn=0012-7183}}</ref> |
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Finns share more [[Identity by descent|identity-by-descent]] (IBD) segments with several other Uralic-speaking peoples, including groups like Estonians, the Sami and the geographically distant Komis and Nganasans, than with their Indo-European-speaking neighbours. This is consistent with the idea that the Uralic peoples share common roots to some degree.<ref name=":3" /> |
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[[File:Suomen geneettinen hienorakenne ennen 1950-lukua.JPG|thumb|Finland's fine-scale genetic structure before the 1950s.]] |
[[File:Suomen geneettinen hienorakenne ennen 1950-lukua.JPG|thumb|Finland's fine-scale genetic structure before the 1950s.]] |
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Finnish genes being often described as homogeneous does not mean that there is no regional variation within Finns.<ref name=":10" /> Finns can be roughly divided into Western and Eastern (or Southwestern and Northeastern) Finnish sub-clusters, which in a fine-scale analysis contain more precise clusters that are consistent with traditional dialect areas.<ref>{{Cite web |date=2017 |title=Itä- ja länsisuomalaiset erottaa jo geeneistä – raja noudattaa Pähkinäsaaren rauhan rajaa |url=https://www.paivanlehti.fi/ita-ja-lansisuomalaiset-erottaa-jo-geeneista-raja-noudattaa-pahkinasaaren-rauhan-rajaa/ |access-date=1 August 2023 |website=paivanlehti.fi |language=fi |archive-date=1 August 2023 |archive-url=https://web.archive.org/web/20230801200216/https://www.paivanlehti.fi/ita-ja-lansisuomalaiset-erottaa-jo-geeneista-raja-noudattaa-pahkinasaaren-rauhan-rajaa/ |url-status=live }}</ref><ref name=":10" /> When looking at the [[fixation index]] (F<sub>ST</sub>) values, the distance within Finns from different parts of the same country is exceptional in Europe. The distance between Western and Eastern Finns is higher than the distance between many European groups from different countries, such as the British people and Northern Germans.<ref name=":4" /><ref>{{cite web |last=Häkkinen |first=Jaakko |date=2011 |title=Seven Finnish populations: the greatest intranational substructure in Europe |url=http://www.mv.helsinki.fi/home/jphakkin/SevenFinnish.xps |archive-url=https://web.archive.org/web/20110927045203/http://www.mv.helsinki.fi/home/jphakkin/SevenFinnish.xps |archive-date=27 September 2011 |access-date=17 August 2011}}</ref><ref name=":10" /> This is also noticeable in the distances of Finns from other Europeans, as the isolation is even more profound in Eastern Finns than in Western Finns.<ref name="Salmela2011" /> The division is related to the later settlement of Eastern Finland by a small number of Finns, who then experienced separate founder and bottleneck effects and genetic drift.<ref name=":10" /><ref name=":6">{{Cite journal |last1=Salmela |first1=Elina |last2=Lappalainen |first2=Tuuli |last3=Fransson |first3=Ingegerd |last4=Andersen |first4=Peter M. |last5=Dahlman-Wright |first5=Karin |last6=Fiebig |first6=Andreas |last7=Sistonen |first7=Pertti |last8=Savontaus |first8=Marja-Liisa |last9=Schreiber |first9=Stefan |last10=Kere |first10=Juha |last11=Lahermo |first11=Päivi |date=24 October 2008 |title=Genome-Wide Analysis of Single Nucleotide Polymorphisms Uncovers Population Structure in Northern Europe |journal=PLOS ONE |language=en |volume=3 |issue=10 |pages=e3519 |bibcode=2008PLoSO...3.3519S |doi=10.1371/journal.pone.0003519 |issn=1932-6203 |pmc=2567036 |pmid=18949038 |doi-access=free}}</ref> Eastern Finns also have a higher portion of autosomal Siberian admixture<ref name=":10" /> and a higher frequency of the Y-haplogroup N1c (71.6%). While N1c is the most common haplogroup in Western Finland as well (53.8%), the haplogroup I1a is found more often there (30.9%) than it is in Eastern and Northern Finland (19%).<ref name=":8">Assessing Finnish Y-chromosomal haplogroups using genotyping array data – Towards understanding the role of Y in complex disease – Annina Preussner 2021 University of Helsinki</ref> This suggests that there is an additional Western component in the Western Finnish gene pool.<ref>Lang, Valter (2020). ''Homo Fennicus – Itämerensuomalaisten etnohistoria''. Finnish Literature Society. p. 94. {{ISBN|978-951-858-130-0}}</ref> Despite the differences, the IBS analysis points out that Western and Eastern Finns share overall a largely similar genetic foundation.<ref name=":6" /><ref>{{Cite web |date=2008 |title=Lehti: Geneettinen analyysi kertoo maakuntasi |url=https://www.iltalehti.fi/uutiset/a/200810248480655 |access-date=22 July 2023 |website=www.iltalehti.fi |language=fi |archive-date=22 July 2023 |archive-url=https://web.archive.org/web/20230722185611/https://www.iltalehti.fi/uutiset/a/200810248480655 |url-status=live }}</ref> |
Finnish genes being often described as homogeneous does not mean that there is no regional variation within Finns.<ref name=":10" /> Finns can be roughly divided into Western and Eastern (or Southwestern and Northeastern) Finnish sub-clusters, which in a fine-scale analysis contain more precise clusters that are consistent with traditional dialect areas.<ref>{{Cite web |date=2017 |title=Itä- ja länsisuomalaiset erottaa jo geeneistä – raja noudattaa Pähkinäsaaren rauhan rajaa |url=https://www.paivanlehti.fi/ita-ja-lansisuomalaiset-erottaa-jo-geeneista-raja-noudattaa-pahkinasaaren-rauhan-rajaa/ |access-date=1 August 2023 |website=paivanlehti.fi |language=fi |archive-date=1 August 2023 |archive-url=https://web.archive.org/web/20230801200216/https://www.paivanlehti.fi/ita-ja-lansisuomalaiset-erottaa-jo-geeneista-raja-noudattaa-pahkinasaaren-rauhan-rajaa/ |url-status=live }}</ref><ref name=":10" /> When looking at the [[fixation index]] (F<sub>ST</sub>) values, the distance within Finns from different parts of the same country is exceptional in Europe. The distance between Western and Eastern Finns is higher than the distance between many European groups from different countries, such as the British people and Northern Germans.<ref name=":4" /><ref>{{cite web |last=Häkkinen |first=Jaakko |date=2011 |title=Seven Finnish populations: the greatest intranational substructure in Europe |url=http://www.mv.helsinki.fi/home/jphakkin/SevenFinnish.xps |archive-url=https://web.archive.org/web/20110927045203/http://www.mv.helsinki.fi/home/jphakkin/SevenFinnish.xps |archive-date=27 September 2011 |access-date=17 August 2011}}</ref><ref name=":10" /> This is also noticeable in the distances of Finns from other Europeans, as the isolation is even more profound in Eastern Finns than in Western Finns.<ref name="Salmela2011" /> The division is related to the later settlement of Eastern Finland by a small number of Finns, who then experienced separate founder and bottleneck effects and genetic drift.<ref name=":10" /><ref name=":6">{{Cite journal |last1=Salmela |first1=Elina |last2=Lappalainen |first2=Tuuli |last3=Fransson |first3=Ingegerd |last4=Andersen |first4=Peter M. |last5=Dahlman-Wright |first5=Karin |last6=Fiebig |first6=Andreas |last7=Sistonen |first7=Pertti |last8=Savontaus |first8=Marja-Liisa |last9=Schreiber |first9=Stefan |last10=Kere |first10=Juha |last11=Lahermo |first11=Päivi |date=24 October 2008 |title=Genome-Wide Analysis of Single Nucleotide Polymorphisms Uncovers Population Structure in Northern Europe |journal=PLOS ONE |language=en |volume=3 |issue=10 |pages=e3519 |bibcode=2008PLoSO...3.3519S |doi=10.1371/journal.pone.0003519 |issn=1932-6203 |pmc=2567036 |pmid=18949038 |doi-access=free}}</ref> Eastern Finns also have a higher portion of autosomal Siberian admixture<ref name=":10" /> and a higher frequency of the Y-haplogroup N1c (71.6%). While N1c is the most common haplogroup in Western Finland as well (53.8%), the haplogroup I1a is found more often there (30.9%) than it is in Eastern and Northern Finland (19%).<ref name=":8">Assessing Finnish Y-chromosomal haplogroups using genotyping array data – Towards understanding the role of Y in complex disease – Annina Preussner 2021 University of Helsinki</ref> This suggests that there is an additional Western component in the Western Finnish gene pool.<ref>Lang, Valter (2020). ''Homo Fennicus – Itämerensuomalaisten etnohistoria''. Finnish Literature Society. p. 94. {{ISBN|978-951-858-130-0}}</ref> Despite the differences, the IBS analysis points out that Western and Eastern Finns share overall a largely similar genetic foundation.<ref name=":6" /><ref>{{Cite web |date=2008 |title=Lehti: Geneettinen analyysi kertoo maakuntasi |url=https://www.iltalehti.fi/uutiset/a/200810248480655 |access-date=22 July 2023 |website=www.iltalehti.fi |language=fi |archive-date=22 July 2023 |archive-url=https://web.archive.org/web/20230722185611/https://www.iltalehti.fi/uutiset/a/200810248480655 |url-status=live }}</ref> |
Revision as of 15:28, 21 November 2024
Total population | |||||||||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
c. 6–7 million[a] | |||||||||||||||||||||||||||||||||||
Regions with significant populations | |||||||||||||||||||||||||||||||||||
Finland c. 4.7–5.1 million[1][2][3][4][b] Other significant population centers: | |||||||||||||||||||||||||||||||||||
United States | 653,222[5] | ||||||||||||||||||||||||||||||||||
Sweden | 156,045[6][c]–712,000[7][d] (including Tornedalians) | ||||||||||||||||||||||||||||||||||
Canada | 143,645[8] | ||||||||||||||||||||||||||||||||||
Russia | 127,600 (with all Karelians)[a][9] 34,300 (with Ingrian Finns) | ||||||||||||||||||||||||||||||||||
Australia | 7,939[10] | ||||||||||||||||||||||||||||||||||
Norway | 15,000–60,000 (including Forest Finns and Kvens)[11][12] | ||||||||||||||||||||||||||||||||||
Germany | 33,000 (2022)[13] | ||||||||||||||||||||||||||||||||||
United Kingdom | 15,000–30,000[14] | ||||||||||||||||||||||||||||||||||
Spain | 17,433 (in 2022)[15] (up to 40,000 part-year residents)[16] | ||||||||||||||||||||||||||||||||||
Estonia | 8,260[17] | ||||||||||||||||||||||||||||||||||
France | 7,000[18] | ||||||||||||||||||||||||||||||||||
Netherlands | 5,000[19] | ||||||||||||||||||||||||||||||||||
Italy | 4,000[20] | ||||||||||||||||||||||||||||||||||
Switzerland | 3,800[21] | ||||||||||||||||||||||||||||||||||
Brazil | 3,100[22] | ||||||||||||||||||||||||||||||||||
Denmark | 3,000[23] | ||||||||||||||||||||||||||||||||||
Belgium | 3,000[24] | ||||||||||||||||||||||||||||||||||
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Languages | |||||||||||||||||||||||||||||||||||
Finnish and its dialects | |||||||||||||||||||||||||||||||||||
Religion | |||||||||||||||||||||||||||||||||||
Predominantly Lutheranism or irreligious, Eastern Orthodox minority[40] | |||||||||||||||||||||||||||||||||||
Related ethnic groups | |||||||||||||||||||||||||||||||||||
Sámi, Balts, and other Baltic Finns Especially Karelians, Izhorians, Vepsians, Kvens, and Tornedalians | |||||||||||||||||||||||||||||||||||
a The total figure is merely a sum of all the referenced populations listed. b No official statistics are kept on ethnicity. However, statistics of the Finnish population according to first language and citizenship are documented and available. |
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Finns or Finnish people (Finnish: suomalaiset, IPA: [ˈsuo̯mɑlɑi̯set]) are a Baltic Finnic[41] ethnic group native to Finland.[42] Finns are traditionally divided into smaller regional groups that span several countries adjacent to Finland, both those who are native to these countries as well as those who have resettled. Some of these may be classified as separate ethnic groups, rather than subgroups of Finns. These include the Kvens and Forest Finns in Norway, the Tornedalians in Sweden, and the Ingrian Finns in Russia.
Finnish, the language spoken by Finns, is closely related to other Balto-Finnic languages, e.g. Estonian and Karelian. The Finnic languages are a subgroup of the larger Uralic family of languages, which also includes Hungarian. These languages are markedly different from most other languages spoken in Europe, which belong to the Indo-European family of languages. Native Finns can also be divided according to dialect into subgroups sometimes called heimo (lit. 'tribe'), although such divisions have become less important due to internal migration.
Today, there are approximately 6–7 million ethnic Finns and their descendants worldwide, with the majority of them living in their native Finland and the surrounding countries, namely Sweden, Russia and Norway. An overseas Finnish diaspora has long been established in the countries of the Americas and Oceania, with the population of primarily immigrant background, namely Australia, Canada, New Zealand, Brazil, and the United States.
Terminology
The Finnish term for Finns is suomalaiset (sing. suomalainen).
It is a matter of debate how best to designate the Finnish-speakers of Sweden, all of whom have migrated to Sweden from Finland. Terms used include Sweden Finns and Finnish Swedes, with a distinction almost always made between more recent Finnish immigrants, most of whom have arrived after World War II, and Tornedalians, who have lived along what is now the Swedish-Finnish border since the 15th century.[43] The term "Finn" occasionally also has the meaning "a member of a people speaking Finnish or a Finnic language".
Etymology
Historical references to Northern Europe are scarce, and the names given to its peoples and geographic regions are obscure; therefore, the etymologies of the names are questionable. Such names as Fenni, Phinnoi, Finnum, and Skrithfinni / Scridefinnum appear in a few written texts starting from about two millennia ago in association with peoples located in a northern part of Europe, but the real meaning of these terms is debatable. It has been suggested that this non-Uralic ethnonym is of Germanic language origin and related to such words as Old High German: finthan, lit. 'find, notice'; Old English: fanthian, lit. 'check, try'; and fendo (Middle High German: vende) 'pedestrian, wanderer'.[44] Another etymological interpretation associates this ethnonym with fen in a more toponymical approach. Yet another theory postulates that the words Finn and Kven are cognates. The Icelandic Eddas and Norse sagas (11th to 14th centuries), some of the oldest written sources probably originating from the closest proximity, use words like finnr and Finnas inconsistently. However, most of the time they seem to mean northern dwellers with a mobile life style. Current linguistic research supports the hypothesis of an etymological link between the Finnish and the Sami languages and other modern Uralic languages. It also supports the hypothesis of a common etymological origin of the toponyms Sápmi (Northern Sami for 'Lapland') and Suomi (Finnish for 'Finland') and the Finnish and Sami names for the Finnish and Sami languages (suomi and saame). Current research has disproved older hypotheses about connections with the names Häme (Finnish for 'Tavastia')[44] and Proto-Baltic *žeme / Slavic землꙗ (zemlja) meaning 'land'.[44][45] This research also supports the earlier hypothesis that the designation Suomi started out as the designation for Southwest Finland (Finland Proper, Varsinais-Suomi) and later for their language and later for the whole area of modern Finland. But it is not known how, why, and when this occurred. Petri Kallio had suggested that the name Suomi may bear even earlier Indo-European echoes with the original meaning of either "land" or "human",[46] but he has since disproved his hypothesis.[45]
The first known mention of Finns is in the Old English poem Widsith which was compiled in the 10th century, though its contents are believed to be older. Among the first written sources possibly designating western Finland as the land of Finns are also two rune stones. One of these is in Söderby, Sweden, with the inscription finlont (U 582), and the other is in Gotland, a Swedish island in the Baltic Sea, with the inscription finlandi (G 319 M) dating from the 11th century.[47]
History
Origins
As other Western Uralic and Baltic Finnic peoples, Finns originated between the Volga, Oka, and Kama rivers in what is now Russia. The genetic basis of future Finns also emerged in this area.[49] There have been at least two noticeable waves of migration to the west by the ancestors of Finns. They began to move upstream of the Dnieper and from there to the upper reaches of the Daugava, from where they eventually moved along the river towards the Baltic Sea in 1250–1000 BC. The second wave of migration brought the main group of ancestors of Finns from the Baltic Sea to the southwest coast of Finland in the 8th century BC.[50][51]
During the 80–100 generations of the migration, Finnish language changed its form, although it retained its Finno-Ugric roots. Material culture also changed during the transition, although the Baltic Finnic culture that formed on the shores of the Baltic Sea constantly retained its roots in a way that distinguished it from its neighbors.[50][51]
Finnish material culture became independent of the wider Baltic Finnic culture in the 6th and 7th centuries, and by the turn of the 8th century the culture of metal objects that had prevailed in Finland had developed in its own way.[50][52] The same era can be considered to be broadly the date of the birth of the independent Finnish language, although its prehistory, like other Baltic Finnic languages, extends far into the past.[52]
Language
Just as uncertain are the possible mediators and the timelines for the development of the Uralic majority language of the Finns. On the basis of comparative linguistics, it has been suggested that the separation of the Finnic and the Sami languages took place during the 2nd millennium BC, and that the Proto-Uralic roots of the entire language group date from about the 6th to the 8th millennium BC. When the Uralic languages were first spoken in the area of contemporary Finland is debated.[citation needed] It is thought that Proto-Finnic (the proto-language of the Finnic languages) was not spoken in modern Finland, because the maximum divergence of the daughter languages occurs in modern-day Estonia. Therefore, Finnish was already a separate language when arriving in Finland. Furthermore, the traditional Finnish lexicon has a large number of words (about one-third) without a known etymology, hinting at the existence of a disappeared Paleo-European language; these include toponyms such as niemi, 'peninsula'.[citation needed] Because the Finnish language itself reached a written form only in the 16th century, little primary data remains of early Finnish life. For example, the origins of such cultural icons as the sauna, and the kantele (an instrument of the zither family) have remained rather obscure.[citation needed]
Livelihood
Agriculture supplemented by fishing and hunting has been the traditional livelihood among Finns. Slash-and-burn agriculture was practiced in the forest-covered east by Eastern Finns up to the 19th century. Agriculture, along with the language, distinguishes Finns from the Sámi, who retained the hunter-gatherer lifestyle longer and moved to coastal fishing and reindeer herding.[citation needed] Following industrialization and modernization of Finland, most Finns were urbanized and employed in modern service and manufacturing occupations, with agriculture becoming a minor employer (see Economy of Finland).
Religion
Christianity spread to Finland from the Medieval times onward and original native traditions of Finnish paganism have become extinct. [citation needed]Finnish paganism combined various layers of Finnic, Norse, Germanic and Baltic paganism. Finnic Jumala was some sort of sky-god and is shared with Estonia. Belief of a thunder-god, Ukko or Perkele, may have Baltic origins.[citation needed] Elements had their own protectors, such as Ahti for waterways and Tapio for forests. Local animistic deities, haltija, which resemble Scandinavian tomte, were also given offerings to, and bear worship was also known.[citation needed] Finnish neopaganism, or suomenusko, attempts to revive these traditions.[citation needed]
Christianity was introduced to Finns from both the west and the east.[54] Swedish kings conquered western parts of Finland in the late 13th century, imposing Roman Catholicism. Reformation in Sweden had the important effect that bishop Mikael Agricola, a student of Martin Luther's, introduced written Finnish, and literacy became common during the 18th century. When Finland became independent, it was overwhelmingly Lutheran Protestant. A small number of Eastern Orthodox Finns were also included, thus the Finnish government recognized both religions as "national religions". In 2017 70.9% of the population of Finland belonged to the Evangelical Lutheran Church of Finland, 1.1% to the Finnish Orthodox Church, 1.6% to other religious groups and 26.3% had no religious affiliation[citation needed]. Whereas, in Russian Ingria, there were both Lutheran and Orthodox Finns; the former were identified as Ingrian Finns while the latter were considered Izhorians or Karelians[citation needed].
Subdivisions
Finns are traditionally assumed to originate from two different populations speaking different dialects of Proto-Finnic (kantasuomi). Thus, a division into Western Finnish and Eastern Finnish is made. Further, there are subgroups, traditionally called heimo,[55][56] according to dialects and local culture. Although ostensibly based on late Iron Age settlement patterns, the heimos have been constructed according to dialect during the rise of nationalism in the 19th century.
- Western[57]
- Southwest Finland and Satakunta: Finns proper (varsinaissuomalaiset)
- Tavastia: Tavastians (hämäläiset)
- Ostrobothnia: Ostrobothnians (pohjalaiset)
- Southern Ostrobothnians (eteläpohjalaiset)
- Central Ostrobothnians (keskipohjalaiset)
- Northern Ostrobothnians (pohjoispohjalaiset)
- Lapland: Lapland Finns (lappilaiset)
- Eastern
- Karelia: Finnish Karelians (karjalaiset); Karelian dialects of Finnish are distinct from the Karelian language spoken in Russia, and most of North Karelia actually speak Savonian dialects
- Savonia: Savonians (savolaiset)
- Kainuu: Kainuu Finns (kainuulaiset)
- Finnish minority groups outside Finland
- Tornedalians (länsipohjalaiset) of Norrbotten, Sweden
- Forest Finns (metsäsuomalaiset) of Sweden and Norway
- Kvens (kveenit) of Finnmark, Norway
- Ingrian Finns (inkerinsuomalaiset) of Ingria, Russia
- Finnish diaspora (ulkosuomalaiset)
- Sweden Finns (ruotsinsuomalaiset), Finnish minority in Sweden
The historical provinces of Finland can be seen to approximate some of these divisions. The regions of Finland, another remnant of a past governing system, can be seen to reflect a further manifestation of a local identity.
Journalist Ilkka Malmberg toured Finland in 1984 and looked into people's traditional and contemporary understanding of the heimos, listing them as follows: Tavastians (hämäläiset), Ostrobothnians (pohjalaiset), Lapland Finns (lappilaiset), Finns proper (varsinaissuomalaiset), Savonians (savolaiset), Kainuu Finns (kainuulaiset), and Finnish Karelians (karjalaiset).[58]
Today the importance of the tribal (heimo) identity generally depends on the region. It is strongest among the Karelians, Savonians and South Ostrobothnians.[59]
The Sweden Finns are either native to Sweden or have emigrated from Finland to Sweden. An estimated 450,000 first- or second-generation immigrants from Finland live in Sweden, of which approximately half speak Finnish. The majority moved from Finland to Sweden following the Second World War, contributing and taking advantage of the rapidly expanding Swedish economy. This emigration peaked in 1970 and has been declining since. There is also Meänkieli, a language developed in partial isolation from standard Finnish, spoken by three minorities, Tornedalians, Kvens and Lantalaiset, in the border area of northern Sweden. The Finnish language as well as Meänkieli are recognized as official minority languages in Sweden.
Genetics
The use of mitochondrial "mtDNA" (female lineage) and Y-chromosomal "Y-DNA" (male lineage) DNA-markers in tracing back the history of human populations has been gaining ground in ethnographic studies of Finnish people (e.g. the National Geographic Genographic Project[60] and the Suomi DNA-projekti.) The most common maternal haplogroup among Finns is H, as 41.5% of Finnish women belong to it. One in four carry the haplogroup U5.[61] It is estimated to be the oldest major mtDNA haplogroup in Europe and is found in the whole of Europe at a low frequency, but seems to be found in significantly higher levels among Finns, Estonians and the Sami people.[60] The older population of European hunter-gatherers that lived across large parts of Europe before the early farmers appeared are outside the genetic variation of modern populations, but most similar to Finns.[62]
With regard to the Y-chromosome, the most common haplogroups of the Finns is N1c, as it is carried by 58–64 percent of Finnish men.[63] N1c, which is found mainly in a few countries in Europe (Estonia, Finland, Latvia, Lithuania, and Russia), is a subgroup of the haplogroup N distributed across northern Eurasia and suggested to have entered Europe from Siberia.[64] The haplogroup N is typical for Uralic-speaking peoples.[61][65] Other Y-DNA haplogroups among Finns include I1a (25 %), R1a (4.3 %), and R1b (3.5 %).[63]
Finns are genetically closest to Karelians, a fellow Balto-Finnic group.[66][61] Finns and Karelians form a cluster with another Balto-Finnic people, the Veps.[67][61][68] They show relative affinity to Northern Russians as well,[69][67][70] who are known to be at least partially descended from Finno-Ugric-speakers who inhabited Northwestern Russia before the Slavs.[61][71]
When not compared to these groups, Finns have been found to cluster apart from their neighboring populations, forming outlier clusters.[73] They are shifted away from the cline that most Europeans belong to[74] towards geographically distant Uralic-speakers like the Udmurts and Mari (while remaining genetically distant from them as well).[75] Their distance from Western Europeans is about the same as their distance from Komis.[69] The Balto-Finnic Estonians are among the genetically closest populations of Finns, but they are less isolated from the European cluster than Finns. Swedes, while being distinct from the Finns, are also closer to Finns than most European populations.[72][76][77]
Finns being an outlier population has to do with their gene pool having reduced diversity[73] and differences in admixture, including Asian influence, compared to most Europeans.[73][65] In general, Europeans can be modelled to have three ancestral components (hunter-gatherer, farmer and steppe), but this model does not work as such for some northeastern European populations, like the Finns and the Sami.[78] While their genome is still mostly European, they also have some additional East Eurasian ancestry (varies from 5 up to 10[79]–13[80] % in Finns). This component is most likely Siberian-related, best represented by the north Siberian Nganasans. The specific Siberian-like ancestry is suggested to have arrived in Northern Europe during the early Iron Age, linked to the arrival of Uralic languages.[61][78] Finns have high steppe or Corded Ware culture-like admixture, and they have less farmer-related and more hunter-gatherer-related admixture than Scandinavians and West and Central Europeans.[65]
Finns share more identity-by-descent (IBD) segments with several other Uralic-speaking peoples, including groups like Estonians, the Sami and the geographically distant Komis and Nganasans, than with their Indo-European-speaking neighbours. This is consistent with the idea that the Uralic peoples share common roots to some degree.[61]
Finnish genes being often described as homogeneous does not mean that there is no regional variation within Finns.[65] Finns can be roughly divided into Western and Eastern (or Southwestern and Northeastern) Finnish sub-clusters, which in a fine-scale analysis contain more precise clusters that are consistent with traditional dialect areas.[81][65] When looking at the fixation index (FST) values, the distance within Finns from different parts of the same country is exceptional in Europe. The distance between Western and Eastern Finns is higher than the distance between many European groups from different countries, such as the British people and Northern Germans.[72][82][65] This is also noticeable in the distances of Finns from other Europeans, as the isolation is even more profound in Eastern Finns than in Western Finns.[76] The division is related to the later settlement of Eastern Finland by a small number of Finns, who then experienced separate founder and bottleneck effects and genetic drift.[65][83] Eastern Finns also have a higher portion of autosomal Siberian admixture[65] and a higher frequency of the Y-haplogroup N1c (71.6%). While N1c is the most common haplogroup in Western Finland as well (53.8%), the haplogroup I1a is found more often there (30.9%) than it is in Eastern and Northern Finland (19%).[84] This suggests that there is an additional Western component in the Western Finnish gene pool.[85] Despite the differences, the IBS analysis points out that Western and Eastern Finns share overall a largely similar genetic foundation.[83][86]
Theories of the origins of Finns
In the 19th century, the Finnish researcher Matthias Castrén prevailed with the theory that "the original home of Finns" was in west-central Siberia.[87]
Until the 1970s, most linguists believed that Finns arrived in Finland as late as the first century AD. However, accumulating archaeological data suggests that the area of contemporary Finland had been inhabited continuously since the end of the ice age, contrary to the earlier idea that the area had experienced long uninhabited intervals. The hunter-gatherer Sámi were pushed into the more remote northern regions.[88]
A hugely controversial theory is so-called refugia. This was proposed in the 1990s by Kalevi Wiik, a professor emeritus of phonetics at the University of Turku. According to this theory, Finno-Ugric speakers spread north as the Ice age ended. They populated central and northern Europe, while Basque speakers populated western Europe. As agriculture spread from the southeast into Europe, the Indo-European languages spread among the hunter-gatherers. In this process, both the hunter-gatherers speaking Finno-Ugric and those speaking Basque learned how to cultivate land and became Indo-Europeanized. According to Wiik, this is how the Celtic, Germanic, Slavic, and Baltic languages were formed. The linguistic ancestors of modern Finns did not switch their language due to their isolated location.[89] The main supporters of Wiik's theory are Professor Ago Künnap of the University of Tartu, Professor Kyösti Julku of the University of Oulu and Associate Professor Angela Marcantonio of the University of Rome. Wiik has not presented his theories in peer-reviewed scientific publications. Many scholars in Finno-Ugrian studies have strongly criticized the theory. Professor Raimo Anttila, Petri Kallio and brothers Ante and Aslak Aikio have rejected Wiik's theory with strong words, hinting strongly to pseudoscience, and even alt-right political biases among Wiik's supporters.[88][90] Moreover, some dismissed the entire idea of refugia, due to the existence even today of arctic and subarctic peoples. The most heated debate took place in the Finnish journal Kaltio during autumn 2002. Since then, the debate has calmed, each side retaining their positions.[91] Genotype analyses across the greater European genetic landscape have provided some credibility to the theory of the Last Glacial Maximum refugia.[92][93][94][95][96] But this does not in any way corroborate or prove that these 'refugia' spoke Uralic/Finnic, as it belies wholly independent variables that are not necessarily coeval (i.e. language spreads and genetic expansions can occur independently, at different times and in different directions).
See also
- Demographics of Finland
- Finnic (disambiguation)
- Finnish (disambiguation)
- Finnish language
- Finnish Americans
- Finnish Brazilians
- Finnish Canadians
- Finnish Australians
- Finnish immigration to North America
- List of Finns
Explanatory notes
- ^ East Karelians are generally considered to be a closely related but separate ethnic group from Finns, rather than a regional subgroup. Not only because of their Eastern Orthodox faith, but also because of their language and ethnic identity.
References
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- ^ "Finn noun" The Oxford Dictionary of English (revised edition). Ed. Catherine Soanes and Angus Stevenson. Oxford University Press, 2005. Oxford Reference Online. Oxford University Press. Tampere University of Technology. 3 August 2007 [1] Archived 9 March 2020 at the Wayback Machine
- ^ "Traditionally, immigrants were described in English and most other languages by an adjective indicating the new country of residence and a noun indicating their country of origin or their ethnic group. The term "Sweden Finns" corresponds to this naming method. Immigrants to the U.S. have, however, always been designated the "other way around" by an adjective indicating the ethnic or national origin and a noun indicating the new country of residence, for example "Finnish Americans" (never "American Finns"). The term "Finnish Swedes" corresponds to this more modern naming method that is increasingly used in most countries and languages because it emphasises the status as full and equal citizens of the new country while providing information about cultural roots. (For more information about these different naming methods see Swedish-speaking Finns.) Other possible modern terms are "Finnish ethnic minority in Sweden" and "Finnish immigrants". These may be preferable because they make a clear distinction between these two very different population groups for which use of a single term is questionable and because "Finnish Swedes" is often used like "Finland Swedes" to mean "Swedish-speaking Finns". It should perhaps also be pointed out that many Finnish and Swedish speakers are unaware that the English word "Finn" elsewhere than in this article usually means "a native or inhabitant of Finland" ("Finn". American Heritage Dictionary. 2000. Archived from the original on 27 October 2007 – via Bartleby.com.
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- ^ Sedergren, J (2002) Evakko – elokuva ja romaani karjalaispakolaisista"Ennen & nyt 3/02, Jari Sedergren: Evakko - elokuva ja romaani pakolaisuudesta". Archived from the original on 24 September 2015. Retrieved 13 January 2008.
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: CS1 maint: bot: original URL status unknown (link). Ennen & nyt 3/2002. Retrieved 13 January 2008. (in Finnish) The reference is a movie review, which however discusses the cultural phenomenon of the evacuation of Finnish Karelia using and analyzing the heimo concept rather generally. - ^ Topelius, Z. (1876) Maamme kirja. Lukukirja alimmaisille oppilaitoksille Suomessa. Toinen jakso. Suom. Johannes Bäckvall. Archived 4 March 2016 at the Wayback Machine (in Finnish) Retrieved 13 January 2008. (in Finnish) Pp. 187 onwards shows the stereotypical generalizations of the heimos listed here.
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This model, however, does not fit well for present-day populations from north-eastern Europe such as Saami, Russians, Mordovians, Chuvash, Estonians, Hungarians, and Finns: they carry additional ancestry seen as increased allele sharing with modern East Asian populations1,3,9,10. Additionally, within the Bolshoy population, we observe the derived allele of rs3827760 in the EDAR gene, which is found in near-fixation in East Asian and Native American populations today, but is extremely rare elsewhere37, and has been linked to phenotypes related to tooth shape38 and hair morphology39 (Supplementary Data 2). To further test differential relatedness with Nganasan in European populations and in the ancient individuals in this study, we calculated f4(Mbuti, Nganasan; Lithuanian, Test) (Fig. 3). Consistent with f3-statistics above, all the ancient individuals and modern Finns, Saami, Mordovians and Russians show excess allele sharing with Nganasan when used as Test populations.
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