Kentrosaurus: Difference between revisions

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[[File:Fossil Kentrosaurus aethiopicus in Museum für Naturkunde Berlin 002.JPG|thumb|Skull and neck of the Berlin mount]]
In the [[mandible]] (lower jaw), only an incomplete right dentary is known from ''Kentrosaurus''.<ref>Hennig, E. (1936). Ein dentale von Kentrurosaurus aethiopicus Hennig. ''Palaeontographica-Supplementbände'', 309-312.</ref> The deep dentary is almost identical in shape to that of ''Stegosaurus'', albeit much smaller. Similarly, the tooth is a typical stegosaurian tooth, small with a widened base and vertical grooves creating five ridges. The dentary has 13 preserved [[Dental alveolus|alveoli]] on the dorsomedial side and it they are slightly convex in lateral and dorsal views. On the surface adjacent to the alveoli, there is a shallow groove bearing small [[Vertebra|foramina]] (small openings in bone) that is similar to grooves on the dentary of the [[Cretaceous]] neornithischian ''[[Hypsilophodon]]'', with one foramina per tooth position. Stegosaurian teeth were small, triangular, and flat; wear facets show that they did grind their food.<ref name="FasWeis04">{{cite book |title=The Evolution and Extinction of the Dinosaurs |vauthors=Fastovsky DE, Weishampel DB |publisher=Cambridge University Press |year=2005 |isbn=978-0-521-81172-9 |veditors=Fastovsky DE, Weishampel DB |edition=2nd |pages=107–30 |chapter=Stegosauria: Hot Plates}}</ref> A single complete cheek tooth is preserved, with a large crown and long root. The crown notably has fewer marginal [[Denticle (tooth feature)|denticles]] and a prominent [[Cingulum (tooth)|cingulum]] compared to ''Stegosaurus'', ''Tuojiangosaurus'', and ''Huayangosaurus''.<ref name=":2" />
 
===Postcrania===
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The vast majority of stegosaurian dinosaurs thus far recovered belong to the [[Stegosauridae]], which lived in the later part of the Jurassic and early Cretaceous, and which were defined by [[Paul Sereno]] as all stegosaurians more closely related to ''Stegosaurus'' than to ''Huayangosaurus''.<ref>Sereno, P.C., 1998, "A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria", ''Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen'' '''210''': 41-83</ref> This group is widespread, with members across the Northern Hemisphere, [[Africa]] and possibly [[South America]].<ref name=":0">{{Cite journal |last1=Rauhut |first1=Oliver W. M. |last2=Carballido |first2=José Luis |last3=Pol |first3=Diego |date=2020-12-10 |title=First osteological record of a stegosaur (Dinosauria, Ornithischia) from the Upper Jurassic of South America |url=https://www.tandfonline.com/doi/full/10.1080/02724634.2020.1862133 |journal=Journal of Vertebrate Paleontology |language=en |volume=40 |issue=6 |pages=e1862133 |doi=10.1080/02724634.2020.1862133 |bibcode=2020JVPal..40E2133R |s2cid=234161169 |issn=0272-4634}}</ref> The South American remains come from [[Chubut Province|Chubut]], [[Argentina]] and consist only of a partial humerus, but the anatomy of the humerus is very similar to that of ''Kentrosaurus'' and both date to the Late Jurassic. In a phylogenetic analysis, the Chubut stegosaurid was recovered in polytomy with ''Kentrosaurus'' as basal stegosaurids, further suggesting that they are closely related.<ref name=":0" />
 
In Hennig's 1915 description, ''Kentrosaurus'' was assigned to the family [[Stegosauridae]] due to the preservation of dermal armor and features like posterodorsally angled neural spines on the caudal vertebrae.<ref name="Hennig1915" /> This is confirmed by modern [[cladistic]] analyses, although in 1915 Stegosauridae was a far more inclusive concept that included some taxa now classified as ankylosaurs. A consecutive narrowing down of this concept caused ''Kentrosaurus'', until the 1980s to be seen as a typical "primitive" stegosaurian,<ref name="Norman1985" /> to be placed in a more derived, higher, position in the stegosaur evolutionary tree. However, recent analyses have consistently found ''Kentrosaurus'' to be in Stegosauridae, though typically as one of the most basal genera in the family.<ref>{{Cite journal |last1=Hui |first1=Dai |last2=Ning |first2=Li |last3=Maidment |first3=Susannah C. R. |last4=Guangbiao |first4=Wei |last5=Yuxuan |first5=Zhou |last6=Xufeng |first6=Hu |last7=Qingyu |first7=Ma |last8=Xunqian |first8=Wang |last9=Haiqian |first9=Hu |last10=Guangzhao |first10=Peng |date=2022-03-30 |title=New stegosaurs from the Middle Jurassic Lower Member of the Shaximiao Formation of Chongqing, China |journal=Journal of Vertebrate Paleontology |volume=41 |issue=5 |pages=e1995737 |doi=10.1080/02724634.2021.1995737 |s2cid=247267743 |issn=0272-4634|doi-access=free }}</ref><ref name=":0" /><ref name=":1">{{Cite journal |last1=Raven |first1=Thomas J. |last2=Maidment |first2=Susannah C. R. |date=2017 |title=A new phylogeny of Stegosauria (Dinosauria, Ornithischia) |url=https://onlinelibrary.wiley.com/doi/10.1111/pala.12291 |journal=Palaeontology |language=en |volume=60 |issue=3 |pages=401–408 |doi=10.1111/pala.12291|bibcode=2017Palgy..60..401R |hdl=10044/1/45349 |s2cid=55613546 |hdl-access=free }}</ref> ''Kentrosaurus'' has many traits not seen in other stegosaurids but seen in basal stegosaurians, such as the presence of a parascapular spine and maxillary teeth with only seven denticles at the margin.<ref name="Galton1982" /><ref name="Galton1990" /> Cladogram of Stegosauria below that includes nearly every known stegosaur genus, recovering ''Kentrosaurus'' as a basal stegosaurid:<ref name=":1" />
{{Clade|{{clade
|1={{clade
|1=[[Ankylosauria]]
|label2='''Stegosauria'''
|2={{clade
|label1=[[Huayangosauridae]]
|1={{clade
|1={{clade
|1=''[[Huayangosaurus taibaii]]''
|2=''[[Chungkingosaurus jiangbeiensis]]'' }}
|2={{clade
|1=''[[Tuojiangosaurus multispinus]]''
|2=''[[Paranthodon africanus]]'' }} }}
|label2=[[Stegosauridae]]
|2={{clade
|1=''[[Jiangjunosaurus junggarensis]]''
|2={{clade
|1=''[[Gigantspinosaurus sichuanensis]]''
|2={{clade
|1= '''''Kentrosaurus aethiopicus'''''
|2={{clade
|1=''[[Dacentrurus armatus]]''
|2={{clade
|1=''[[Loricatosaurus priscus]]''
|2={{clade
|1={{clade
|1=''[[Hesperosaurus mjosi]]''
|2=''[[Miragaia longicollum]]'' }}
|2={{clade
|1=''[[Stegosaurus stenops]]''
|2=''[[Wuerhosaurus homheni]]'' }} }} }} }} }} }} }} }} }} }}|style=font-size: 85%; line-height:85%;margin:1em auto;|label1=[[Eurypoda]]}}
 
The [[type species|type]] and sole accepted species of ''Kentrosaurus'' is ''Kentrosaurus aethiopicus'', named by Hennig in 1915. Fragmentary fossil material from Wyoming, named ''Stegosaurus longispinus'' by [[Charles W. Gilmore|Charles Gilmore]] in 1914,<ref name="Gilmore14" /> was in 1993 classified as a North American species of ''Kentrosaurus'', as ''K. longispinus''.<ref>{{cite journal | last1 = Olshevsky | first1 = G. | last2 = Ford | first2 = T.L. | year = 1993 | title = The Origin and Evolution of the Stegosaurs | journal = Kyoryugaku Saizensen | volume = 4 | pages = 64–103 }}</ref> However, this action was not accepted by the paleontological community, and ''S. longispinus'' has been assigned to its own genus, ''[[Alcovasaurus]]'', differing from ''Kentrosaurus'' in having more elongated tail spikes and the structure of the pelvis and vertebrae.<ref>Ulansky, R. E., 2014. Evolution of the stegosaurs (Dinosauria; Ornithischia). Dinologia, 35 pp. [in Russian]. [DOWNLOAD PDF] http://dinoweb.narod.ru/Ulansky_2014_Stegosaurs_evolution.pdf.</ref><ref>Ulansky, RE, 2014. Natronasaurus longispinus, 100 years with another name. Dinologia, 10 pp. [In Russian].</ref> Cladogram of the phylogenetic analysis of Stegosauridae conducted by Maidment ''et al'' (2019), which recovers a distinct ''Alcovasaurus'':<ref>{{Cite journal |last1=Maidment |first1=Susannah C. R. |last2=Raven |first2=Thomas J. |last3=Ouarhache |first3=Driss |last4=Barrett |first4=Paul M. |date=2020-01-01 |title=North Africa's first stegosaur: Implications for Gondwanan thyreophoran dinosaur diversity |url=https://www.sciencedirect.com/science/article/pii/S1342937X19302217 |journal=Gondwana Research |language=en |volume=77 |pages=82–97 |doi=10.1016/j.gr.2019.07.007 |bibcode=2020GondR..77...82M |hdl=10141/622706 |s2cid=202188261 |issn=1342-937X|hdl-access=free }}</ref>
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<ref name=Maier2003>{{cite book|last=Maier|first=G|title=African Dinosaurs Unearthed. The Tendaguru Expeditions|publisher=Indiana University Press|location=Bloomington|page=432|isbn=978-0-253-34214-0|year=2003}}</ref>
<ref name=MallisonRealLecto>{{cite journal|last=Mallison|first=H.|year=2011|title=The real lectotype of ''Kentrosaurus aethiopicus'' HENNIG 1915|journal=Neues Jahrbuch für Geologie und Paläontologie|volume=259|issue=2|pages=197–206|url=http://www.schweizerbart.de/resources/downloads/paper_previews/75591.pdf|doi=10.1127/0077-7749/2011/0114}}</ref>
<ref name=MallROM>{{cite journal|last=Mallison|first=H.|year=2010|title=CAD assessment of the posture and range of motion of ''Kentrosaurus aethiopicus'' HENNIG 1915|journal=[[Swiss Journal of Geosciences]]|volume=103|issue=2|pages=211–233|doi=10.1007/s00015-010-0024-2|s2cid=132746786|doi-access=free|bibcode=2010SwJG..103..211M }}</ref>
<ref name=MallCAE>{{cite journal|last=Mallison|first=H.|year=2011|title=Defense capabilities of ''Kentrosaurus aethiopicus'' HENNIG 1915|url=http://www.palaeo-electronica.org/2011_2/255/index.html|journal=Palaeontologia Electronica|volume=14|issue=2|pages=10}}</ref>
<ref name=Mateus2006>{{cite book|title=Paleontology and Geology of the Upper Morrison Formation|journal=Bulletin of the New Mexico Museum of Natural History and Science|year=2006|chapter=Late Jurassic dinosaurs from the Morrison Formation (USA), the Lourinhā and Alcobaça formations (Portugal), and the Tendaguru Beds (Tanzania): a comparison |editor=Foster, J.R. |editor2=Lucas, S.G.|author=Mateus, Octávio |pages=223–232|publisher=New Mexico Museum of Natural History and Science (bulletin 36)|issn=1524-4156}}</ref>