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{{Short description|Dog-wolf hybrid}}
{{Distinguish|Wolfhound (disambiguation)}}
{{pp-semi-indef}}
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}} <!-- End Infobox -->
A '''wolfdog''' is a [[Canis|canine]] produced by the [[Canine reproduction|mating]] of a domestic [[
==Admixture==
[[File:F1 wolf-dog hybrids from Wildlife Park Kadzidlowo, Poland.png|thumb|Hybrids in eastern Poland in the Wildlife Park [[Kadzidłowo]]. To the left: Parents: female wolf and male [[Gończy Polski]]; right: parents: female wolf and male [[West Siberian Laika]].]]
There are a range of experts who believe that they can tell the difference between a wolf, a dog, and a wolfdog, but they have been proven to be incorrect when providing their evidence before courts of law.<ref name="Pierotti-Fogg-2017-co‑evld"/>
[[Genetic admixture|Admixture]] between domestic dogs and [[Subspecies of Canis lupus|other subspecies of gray wolves]] are the most common wolfdogs since dogs and gray wolves are considered the same species, are genetically very close and have shared vast portions of their ranges for millennia. Such admixture in the wild have been detected in many populations scattered throughout Europe and North America, usually occurring in areas where wolf populations have declined from human impacts and persecutions.<ref name=Randi-Hulva-Fabbri-etal-2014/><ref name=Vila-Wayne-2001-12-24-Hybdzn/>
At the same time, wolfdogs are also often bred in captivity for various purposes.
|title=Great Lakes-boreal wolf
|website=Wolf.nrdpfc.ca
|url=http://wolf.nrdpfc.ca/greatlakeswolf.htm
|access-date=2016-04-06
|archive-date=2020-02-22
|archive-url=https://web.archive.org/web/20200222165757/http://wolf.nrdpfc.ca/greatlakeswolf.htm
|url-status=dead
}}</ref>
At the same time, because many isolated populations of the three wolf species in North America have also mixed with coyotes in the wild,<ref name=Hailer-Leonard-2008-10-08-coywolf/>
it has been speculated by some biologists that some of the [[coywolf|coywolf hybrids]] in the northeastern third of the continent may also have both [[coydog]]s and wolfdogs in their gene pool.<ref>
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|title=Mysteries That Howl and Hunt
|newspaper=[[The New York Times]]
|date=27 September 2010
|url=https://www.nytimes.com/2010/09/28/science/28coyotes.html
|access-date=2016-04-06
|last1=Yoon
|first1=Carol Kaesuk
}}
</ref>
Hybrids between dogs and Ethiopian wolves discovered in the [[Ethiopian Highlands]] likely originated from past interactions between free-roaming feral dogs and Ethiopian wolves living in isolated areas.<ref>
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|title=New world wolves and coyotes owe debt to dogs
|newspaper=[[The New York Times]]
|date=5 February 2009
|url=https://www.nytimes.com/2009/02/06/science/06wolves.html
|access-date=2014-03-04
|last1=Derr
|first1=Mark
}}
</ref>
[[Adolph Murie]] was among the first wolf biologists to speculate that the wide color variation in wolves was due to interbreeding with dogs;<ref>
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</ref>
{{
In 2008, it was discovered that a gene [[mutation]] responsible for the [[protein]] [[beta-defensin 3]] is responsible for the black coat color in dogs.<ref name="solved">
{{cite web
|title=Black Wolf Mystery Solved
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| url = http://www.wolfpark.org/wolfdogs/Poster_section1.html
| access-date = <!-- 2007-05-11 --> 2021-09-10 |url-status=dead
|archive-url = https://web.archive.org/web/20070303000105/http://www.wolfpark.org/wolfdogs/Poster_section1.html <!-- Bot retrieved archive -->
|archive-date = 2007-03-03
}}
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|last1=Wayne |first1=Robert K.
|last2=Ostrander |first2=Elaine A.
|s2cid=5547543
}}
</ref>
In first-generation wolfdogs, [[Wolf|gray wolves]] are most often crossed with wolf-like dogs (such as [[German Shepherd|German Shepherd Dog]]s, [[Siberian Husky|Siberian Huskies]], and [[Alaskan Malamute]]s) for an appearance most appealing to owners desiring an [[exotic pet]].<ref name="DEFRA">{{cite web
| url = http://www.defra.gov.uk/wildlife-countryside/pdf/protection/dwa-wolfdogs.pdf
| title = The Keeping of Wolf-Hybrids in Great Britain
| publisher = [[RSPCA]]
| access-date = 2008-06-11
| archive-date = 2008-12-06
| archive-url = https://web.archive.org/web/20081206062712/http://www.defra.gov.uk/wildlife-countryside/pdf/protection/dwa-wolfdogs.pdf
| url-status = dead
}}</ref>
==Documented breeding==
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[[File:Saarloos.jpg|thumb|left|A Saarloos Wolfdog]]
In 1932, [[Netherlands|Dutch]] breeder [[Leendert Saarloos]] crossed a male [[German Shepherd]] dog with a female [[European wolf]]. He then bred the female offspring back with the male German Shepherd
{{cite web
|url=https://www.ukcdogs.com/saarloosewolfdog
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=== The Czechoslovakian Wolfdog ===
[[File:Czechoslovakian-wolfdog-profile big.jpg|thumb|A [[Czechoslovakian Wolfdog]]]]
In the 1950s, the [[Czechoslovakian Wolfdog]] was also created to work on border patrol in the countries now known as [[Slovakia]] and the [[Czech Republic]]. It was originally bred from lines of
=== Volkosob ===
The Volkosob ({{Langx|ru|Волкособ}}, plural: {{Langx|ru|Волкособы|translit=Volkosoby|label=none}}) was initially developed in the 1990s after the [[Dissolution of the Soviet Union|fall of the Soviet Union]]. Russian [[border guard]]s wanted a dog that would possess the trainability and pack mentality of the German Shepherd, combined with the strength, superior senses and cold-resistance of a wild wolf, able to cope in the harsh conditions of the vast Russian borders. In 2000, a [[Steppe wolf|Caspian Steppe Wolf]], noted for being unusually friendly and cooperative towards humans, was bred with German Shepherds of an [[East European Shepherd]] line, until an F3 generation was standardised. Unlike the previous hybrids, the Volkosob was the only breed that was an effective border guardian as they are renowned for not being too shy.
==Livestock guardian dogs==
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Admixture in the wild usually occurs near human habitations where wolf density is low and dogs are common.<ref name="WOLVES">
{{cite book
|first1=L. David |
|first2=Luigi |last2=Boitani
|title=Wolves: Behaviour, Ecology and Conservation
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|first1=Dmitrij, Prof. Dr. |last1=Iwanowitsch
|year=1990
|title=Der Wolf |
|trans-title=The Wolf <!-- Bibikow Moskau, 2. -->
|publisher=Auflage; A. Ziemsen Verlag
|place=Wittenberg Lutherstadt
|
}}
</ref>
Wild wolfdogs were occasionally hunted by European aristocracy, and were termed ''lycisca'' to distinguish them from common wolves.<ref name=Rousseau-2006-Anth-duLoup/>
Noted historic cases (such as the [[Beast of Gévaudan]]) of large wolves that were abnormally aggressive toward humans, may be attributable to wolf-dog mating.<ref name=Willems-1994-AWIC/>
In Europe, unintentional mating of dogs and wild wolves have been confirmed in some populations through [[genetic testing]]. As the survival of some Continental European wolf packs is severely threatened, scientists fear that the creation of wolfdog populations in the wild is a threat to the continued existence of European wolf populations.<ref name=Rincon-2004-04-08-BBC-N/>
However, extensive admixture between wolf and dog is not supported by morphological evidence, and analyses of [[mtDNA]] sequences have revealed that such mating are rare.<ref name="WOLVES"/>
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}}
</ref>
In Canada, the provinces of Alberta, Manitoba, Newfoundland and Labrador, and Prince Edward Island prohibit wolfdogs as pets.<ref name=Bruch-2007-Almanac/>
Most European nations have either outlawed the animal entirely or put restrictions on ownership.<ref>
{{cite web
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Due to the variability inherent to their admixture,<ref name=Willems-1994-AWIC/> whether a wolf–dog cross should be considered more dangerous than a dog depends on behavior specific to the individual alone rather than to wolfdogs as a group.
The view that aggressive characteristics are inherently a part of wolfdog temperament has been contested in recent years by wolfdog breeders and other advocates of wolfdogs as pets.<ref>{{cite web
|url=http://www.inetdesign.com/wolfdunn/wolfdogfaq/pets.html
|title=The Wolf Dunn's Wolfdog FAQ - Question #1
|publisher=Inetdesign.com
|access-date=2013-12-16
|archive-date=2012-08-19
|archive-url=https://web.archive.org/web/20120819140650/http://www.inetdesign.com/wolfdunn/wolfdogfaq/pets.html
|url-status=dead
}}</ref><ref name="The Wolf Is At The Door, Inc">
{{cite web
|title=The Wolf is at the Door, Inc
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==In popular culture==
* [[Jed (wolfdog)|Jed]] was a Canadian timber wolf-Alaskan Malamute and animal actor, known for his roles in such movies as [[White Fang (1991 film)|''White Fang'']] (1991), ''[[White Fang 2: Myth of the White Wolf]]'' (1994), ''[[The Journey of Natty Gann]]'' (1985), and [[The Thing (1982 film)|''The Thing'']] (1982); he was born in 1977 and died in June 1995, aged 18.
* Balto, Aleu and Kodi are fictitious wolfdogs in the animated films ''[[Balto (film)|Balto]]'' (1995), ''[[Balto II: Wolf Quest]]'' (2002) and ''[[Balto III: Wings of Change]]'' (2004), respectively. The actual [[Balto]] was not a wolfdog but instead a [[Siberian Husky]].
* [[White Fang]] is the titular character of [[Jack London]]'s eponymous [[White Fang|1906 novel]], first serialized in [[Outing (magazine)|''Outing'' magazine]], that details the wild wolfdog's journey to domestication in the [[Yukon Territory]] and the [[Northwest Territories]] during the 1890s [[Klondike Gold Rush]].
* ''[[The Wolf Dog]]'' (1933) is an American [[Pre-code Hollywood|Pre-Code]] Mascot film serial starring Frankie Darro and [[Rin Tin Tin]] Jr.
* ''[[Wolf Dog]]'' (1958), also known as ''A Boy and His Dog'', is a Northwestern movie, directed and produced by Sam Newfield, and produced by Regal Films
* ''Wolfdogs Magazine'' self-describes as a progressive "community based publication for wolfdog enthusiasts".<ref>{{cite book
|url=http://www.wolfdogsmagazine.org/
|title=Wolfdogs Magazine
|access-date=2009-08-23
|archive-date=2013-07-21
|archive-url=https://web.archive.org/web/20130721115110/http://wolfdogsmagazine.org/
|url-status=dead
}}</ref>
==Further reading==
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|author2=Miller, Andrew
|title=Between Dog and Wolf: Understanding the connection and confusion
}}{{full citation needed|date=September 2021|reason=publisher, publ. date, ISBN}}
* {{cite book
|author=Prendergast, Dorothy
|title=Above Reproach: A guide for wolf hybrid owners
}}{{full citation needed|date=September 2021|reason=publisher, publ. date, ISBN}}
* {{cite book
|author=Prendergast, Dorothy
|title=The Wolf Hybrid
}}{{full citation needed|date=September 2021|reason=publisher, publ. date, ISBN}}
* {{cite book
|author=Wilde, Nicole
|title=Living with Wolfdogs
}}{{full citation needed|date=September 2021|reason=publisher, publ. date, ISBN}}
* {{cite book
|author=Wilde, Nicole
|title=Wolfdogs A-Z: Behavior, training & more
}}{{full citation needed|date=September 2021|reason=publisher, publ. date, ISBN}}
==See also==
{{Commons category|Wolf-dog hybrid}}{{Portal|Dogs
}}
* [[Black wolf]]
* [[Canid hybrid]]
* [[Coydog]]
* [[Coywolf]]
* [[Dingo-dog hybrid]]
* [[Dogxim]]
* [[Jackal–dog hybrid]]
* [[Wolves as pets and working animals]]
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</ref>
<ref name=DEFRA-RSPCA-2013-01-23>{{cite web
|url=http://www.defra.gov.uk/wildlife-countryside/gwd/wolfdogs/wolfdogs.pdf
|title=RSPCA report on wolfdogs
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|date=2013-01-23
|access-date=2013-12-16
|archive-date=2006-12-15
|archive-url=https://web.archive.org/web/20061215025632/http://www.defra.gov.uk/wildlife-countryside/gwd/wolfdogs/wolfdogs.pdf
|url-status=dead
}}</ref>
<ref name=Frantz-Larson-2018-ch-1>
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</ref>
<ref name=Ottawa-Humane-c2013>{{cite web
|title=Ottawa Humane
|website=Ottawahumane.ca
|url=http://www.ottawahumane.ca/wolfdoghybrids.html
|access-date=2013-12-16
|archive-date=2013-12-16
|archive-url=https://web.archive.org/web/20131216043033/http://www.ottawahumane.ca/wolfdoghybrids.html
|url-status=dead
}}</ref>
<ref name=Humane-Soc-c2007>
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|pmid=18841199 |pmc=2556088
|bibcode=2008PLoSO...3.3333H
|doi-access=free
}}
</ref>
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|volume=105 |issue=3 |pages=345–353
|doi=10.1093/jhered/esu014 |pmid=24622972
|doi-access=
}}
</ref>
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|archive-date=June 17, 2009
}}
— The archived documents are two open letters, the June 1997 from R. McBryde to D.R. Parsons, and
</ref>
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</ref>
<ref name="Pierotti-Fogg-2017-co‑evld">
{{cite book
|last1=Pierotti |first1=R. |last2=Fogg |first2=B.
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|pmid=24466077 |pmc=3899229
|bibcode=2014PLoSO...986409R
|doi-access=free }}
</ref>
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| page= 319
| isbn= 978-2-603-01335-9
|
}}
</ref>
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|volume=13 |pages=195–198
|journal=[[Conservation Biology (journal)|Conservation Biology]]
|s2cid = 85011290
}}
</ref>
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</ref>
<ref name=Willems-1994-AWIC>{{cite web
|first = Robert A.
|
|title = The wolf-dog hybrid: An overview of a controversial animal
|date = Winter 1994–1995
|orig-date = 1994 |periodical = AWIC Newsletter
|volume
|number = 4
|pages = 3–7
|issn = 1050-561X
|lccn = 90002352
|place = Beltsville, MD
|department = Animal Welfare Information Center / National Agricultural Library
|publisher = [[United States Department of Agriculture]]
|url = http://www.nal.usda.gov/awic/newsletters/v5n4/5n4wille.htm
|
|url-status = dead
|archive-url = https://web.archive.org/web/20141221144425/http://www.nal.usda.gov/awic/newsletters/v5n4/5n4wille.htm
|archive-date = 2014-12-21
}}</ref>
}} <!-- end "refs=" -->
==External links==
* {{cite web|url=http://texx-wolf-tails.webs.com |title=Wolf-dog Education|website=texx-wolf-tails.webs.com}}
* {{cite web|url=http://kyrawolfdog.webs.com |title=Wolfdog Education: the Basics|website=kyrawolfdog.webs.com}}
* {{cite web|url=http://www.renokeo.com/wrr.html
{{Mammal hybrids}}
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