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Proto-Afroasiatic homeland

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The Afroasiatic languages, as they are distributed today

The Proto-Afroasiatic homeland is the hypothetical place where speakers of the Proto-Afroasiatic language lived in a single linguistic community, or complex of communities, before this original language dispersed geographically and divided into separate distinct languages. Afroasiatic languages are today mostly distributed in parts of Africa, and Western Asia.

The contemporary Afroasiatic languages are spoken in West Asia, North Africa, the Horn of Africa, parts of the Sahara and Sahel, and Malta. The various hypotheses for the Afroasiatic homeland are distributed throughout this territory;[1][2][3][4] that is, it is generally assumed that proto-Afroasiatic was spoken in some region where Afroasiatic languages are still spoken today. However, there is disagreement as to which part of the contemporary Afroasiatic speaking areas corresponds with the original homeland. The majority of scholars today contend that Afroasiatic languages arose somewhere in Northeast Africa.[5]

Date of Proto-Afroasiatic

[edit]

There is no consensus as to when Proto-Afroasiatic was spoken.[6] The absolute latest date for when Proto-Afroasiatic could have been extant is c. 4000 BC, after which Egyptian and the Semitic languages are firmly attested. However, in all likelihood these languages began to diverge well before this hard boundary.[7] The estimations offered by scholars as to when Proto-Afroasiatic was spoken vary widely, ranging from 18,000 BC to 8,000 BC. According to Igor M. Diakonoff (1988: 33n), proto-Afroasiatic was spoken c. 10,000 BC. According to Christopher Ehret (2002: 35–36), proto-Afroasiatic was spoken c. 11,000 BC at the latest, and possibly as early as c. 16,000 BC. These dates are older than dates associated with most other proto-languages.[6] An estimate at the youngest end of this range still makes Afroasiatic the oldest proven language family.[8] Contrasting proposals of an early emergence, Tom Güldemann has argued that less time may have been required for the divergence than is usually assumed, as it is possible for a language to rapidly restructure due to areal contact, with the evolution of Chadic (and likely also Omotic) serving as pertinent examples.[9]

Urheimat hypotheses

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No consensus exists as to where proto-Afroasiatic originated. Scholars have proposed locations for the Afroasiatic homeland across Africa and western Asia.[6][10] A complicating factor is the lack of agreement on the subgroupings of Afroasiatic (see Further subdivisions) – this makes associating archaeological evidence with the spread of Afroasiatic particularly difficult.[11] Nevertheless, there is a long-accepted link between the speakers of Proto-Southern Cushitic languages and the East African Savanna Pastoral Neolithic (3000 BC), and archaeological evidence associates the Proto-Cushitic speakers with economic transformations in the Sahara dating c. 8,500 years ago, as well as the speakers of the Proto-Zenati variety of the Berber languages with an expansion across the Maghreb in the 5th century AD.[12] More hypothetical links associate the proto-Afroasiatic-speakers with the Kebaran and the Mushabian culture,[13] while others argue for a possible affiliation between proto-Afroasiatic and the Natufian culture.[6][14][15][16]

The linguistic view on the location of the homeland of Afroasiatic languages is largely divided into proponents for a homeland within Africa, and proponents for a homeland in western Asia. To date, a homeland within Africa is favored by a majority of scholars, although a significant minority of scholars support a homeland in western Asia.[6][7]

Pagani and Crevecoeur (2019) argue that, given the still open debate on the origin of Afroasiatic, the consensus will probably settle on an intermediate "across-the-Sinai" solution. They also note that the very early interactions between African and Eurasian cultures, point "to a geographical shrinking of what can currently be defined as 'strictly African' in a long term perspective."[17]

Western Asian homeland theory

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Levant agriculturalists

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Supporters of a western Asian origin for Afroasiatic are particularly common among those with a background in Semitic or Egyptological studies,[11] and amongst archaeological proponents of the "farming/language dispersal hypothesis" according to which major language groups dispersed with early farming technology in the Neolithic.[18][19] The leading linguistic proponent of this idea in recent times is Alexander Militarev, who argues that Proto-Afroasiatic was spoken by early agriculturalists in the Levant and subsequently spread to Africa. Militarev associates the speakers of Proto-Afroasiatic with the Levantine Post-Natufian Culture, arguing that the reconstructed lexicon of flora and fauna, as well as farming and pastoralist vocabulary indicates that Proto-AA must have been spoken in this area. Scholar Jared Diamond and archaeologist Peter Bellwood have taken up Militarev's arguments as part of their general argument that the spread of linguistic macrofamilies (such as Afroasiatic, Bantu, and Austroasiatic) can be associated with the development of agriculture; they argue that there is clear archaeological support for farming spreading from the Levant into Africa via the Nile valley.[14][16]

Militarev, who linked proto-Afroasiatic to the Levantine Natufian culture, that preceded the spread of farming technology, believes the language family to be about 10,000 years old. He wrote (Militarev 2002, p. 135) that the "Proto-Afrasian language, on the verge of a split into daughter languages", meaning, in his scenario, into "Cushitic, Omotic, Egyptian, Semitic and Chadic-Berber", "should be roughly dated to the ninth millennium BC". Support for the migration of agricultural populations, according to linguists, are the word for dog (an Asian domesticate) reconstructed to Proto-Afroasiatic[20] as well as words for bow and arrow,[21] which according to some archaeologists spread rapidly across North Africa once they were introduced to North Africa from the Near East, viz. Ounan points.[22]

Lexicon linked to a pastoralist society (cattle-breeding) reconstructed for proto-Afroasiatic also support a western Asian homeland, possibly indicating an earlier pastoralist migration.[23]

Northeast African homeland theory

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Afroasiatic languages ca. 500 BC, Omotic and Chadic branches are not shown[24]

A Northeast African homeland has been proposed by the majority of linguists as the origin of the language group because it includes the geographic center of its present distribution and the majority of the diversity observed among the Afroasiatic language family, sometimes considered a telltale sign for a linguistic geographic origin.[5][25] Within this hypothesis there are a number of competing variants:

Red Sea coast

[edit]
Proposed homeland of the Afroasiatic languages and subsequent migration of the Semitic branch into the Levant
African languages and their place of origin according to Christopher Ehret; "Afrasian" is Ehret's preferred term for Afroasiatic.

Christopher Ehret has proposed the western Red Sea coast from Eritrea to southeastern Egypt. While Ehret disputes Militarev's proposal that Proto-Afroasiatic shows signs of a common farming lexicon, he suggests that early Afroasiatic languages were involved in the even earlier development of intensive food collection in the areas of Ethiopia and Sudan. In other words, he proposes an even older age for Afroasiatic than Militarev, at least 11,000 years old, and believes farming lexicon can only be reconstructed for branches of Afroasiatic. Ehret argues that Proto-Afroasiatic speakers in Northeast Africa developed subsistence patterns of intensive plant collection and pastoralism, giving the population an economic advantage which impelled the expansion of the Afroasiatic languages. He suggests that a Proto-Semitic or Proto-Semito-Berber-speaking population migrated from Northeast Africa to the Levant during the late Paleolithic.[26][27][28][29]

In the next phase, unlike many other authors Ehret proposed an initial split between northern, southern and Omotic. The northern group includes Semitic, Egyptian and Berber (agreeing with others such as Diakonoff). He proposed that Chadic stems from Berber (some other authors group it with southern Afroasiatic languages such as Cushitic ones).

Ethiopia

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Roger Blench has proposed a region in the adjacent Horn of Africa, specifically in modern day Ethiopia, arguing that Omotic represents the most basal branch and displays high diversity.[1] Others have however pointed out that Omotic displays strong signs of contact with non-Afroasiatic languages, with some arguing that Omotic should be regarded as an independent language family.[30][31] Like Ehret, Blench accepts that Omotic is part of the Afroasiatic grouping and sees the split of northern languages from Omotic as an important early development. Güldemann (2018) does not accept Omotic as unified group, but argues for at least four distinct groupings.[9]

Sahel/Sahara

[edit]
Sahara and Sahel

Igor Diakonoff proposed the Eastern Saharan region, specifically the southern fringe of the Sahara as possible location of the Afroasiatic homeland.[3][32] Lionel Bender proposed the area near Khartoum, Sudan, at the confluence of the Blue Nile and White Nile.[3][32] The details of his theory are widely cited but controversial, as it involves the proposal that Semitic originated in Ethiopia and crossed to Asia directly from there over the Red Sea.[33]

Evidence from population genetics

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Autosomal DNA

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Scholars, such as Hodgson et al., present archaeogenetic evidence in favor for a place of dispersion within Africa, but argue that the speakers of Proto-Afroasiatic can ultimately be linked to a Paleolithic and pre-agricultural migration wave into Africa from Western Asia, and that the Semitic-branch represents a later back-migration to the Levant.[34]

Pre-Neolithic and Neolithic migration events in Africa.[35]
The "Ethio-Somali" genetic component labelled in green.[36]

According to an autosomal DNA research in 2014 on ancient and modern populations, the Afroasiatic languages likely spread across Africa and the Near East by an ancestral population(s) carrying a newly identified "non-African" (Western Eurasian) genetic component, which the researchers dub the "Ethio-Somali" component. This genetic component is most closely related to the "Maghrebi" component and is believed to have diverged from other "non-African" (Western Eurasian) ancestries at least 23,000 years ago. The "Ethio-Somali" genetic component is prevalent among modern Afroasiatic-speaking populations, and found at its highest levels among Cushitic peoples in the Horn of Africa. On this basis, the researchers suggest that the original Ethio-Somali carrying population(s) probably arrived in the pre-agricultural period (12–23 ka) from the Near East, having crossed over into northeast Africa via the Sinai Peninsula and then split into two, with one branch continuing west across North Africa and the other heading south into the Horn of Africa. They suggest that a descendant population migrated back to the Levant prior to 4000 BC and developed the Semitic branch of Afroasiatic. Later migration from Arabia into the HOA beginning around 3 ka would explain the origin of the Ethio-Semitic languages at this time.[37] A similar view has already been proposed earlier, suggesting that the ancestors of Afroasiatic speakers could have been a population originating in the Near East that migrated to Northeast Africa during the Late Palaeolithic with a subset later moving back to the Near East.[38]

Subsequent archaeogenetic studies have corroborated the migrations of Western Eurasian ancestry during the Paleolithic into Africa, becoming the dominant component of Northern Africa since at least 15,000 BC. The "Maghrebi" component, which gave rise to the Iberomaurusian culture, is described as autochthonous to Northern Africa, related to the Paleolithic Eurasian migration wave, and the characteristic ancestry components of modern Northern Africans along a West-to-East cline, with Northeastern Africans having an additionally higher frequency of a Neolithic Western Asian component associated with the Neolithic expansion.[39]

Genetic research on Afroasiatic-speaking populations revealed strong correlation between the distribution of Afroasiatic languages and the frequency of Northern African/Natufian/Arabian-like ancestry. In contrast, Omotic speakers display ancestry mostly distinct from other Afroasiatic-speakers, indicating language shift, or support for the exclusion of Omotic from the Afroasiatic group.[40]

Genetic studies on a specimen of the Savanna Pastoral Neolithic excavated at the Luxmanda site in Tanzania, which has been associated with migrations of Cushitic-speaking peoples and the spread of pastoralism, found that the specimen carried a large proportion of ancestry related to the Pre-Pottery Neolithic culture of the Levant (Natufian), similar to that borne by modern Afroasiatic-speaking populations inhabiting the Horn of Africa. It is suggested that a population related to the Pre-Pottery Neolithic culture of the Levant contributed significantly to historical Eastern African populations represented by the c. 5,000 year old Luxmanda specimen, while modern Cushitic-speaking populations have additional contributions from Dinka-related and "Neolithic Iranian-related" sources. This type of ancestry was later partially replaced by following migration events associated with the Bantu expansion, with Bantu-speaking Eastern Africans having only little ancestry associated with the Pre-Pottery Neolithic culture of the Levant.[41][42]

Y-chromosome evidence

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Keita (2008) examined a published Y-chromosome dataset on Afro-Asiatic populations and found that a key lineage E-M35/E-M78, sub-clade of haplogroup E, was shared between the populations in the locale of Egyptian and Libyan speakers and modern Cushitic speakers from the Horn. These lineages are present in Egyptians, Berbers, Cushitic speakers from the Horn of Africa, and Semitic speakers in the Near-East. He noted that variants are also found in the Aegean and Balkans, but the origin of the M35 subclade was in Egypt or Libya, and its clades were dominant in a core portion of Afro-Asiatic speaking populations which included Cushitic, Egyptian and Berber groups, in contrast Semitic speakers showed a decline in frequency going west to east in the Levantine-Syria region. Keita identified high frequencies of M35 (>50%) among Omotic populations, but stated that this derived from a small, published sample of 12. Keita also wrote that the PN2 mutation was shared by M35 and M2 lineages and this paternal clade originated from East Africa. He concluded that "the genetic data give population profiles that clearly indicate males of African origin, as opposed to being of Asian or European descent" but acknowledged that the biodiversity does not indicate any specific set of skin colors or facial features as populations were subject to microevolutionary pressures.[43]

Fregel summarized that the Y-chromosome diversity of North Africans was compatible with a demic expansion from the Middle East, because the age of common lineages in North Africa (E-M78 and J-304) were relatively recent. The North African pattern of Y-chromosome variation was mostly shaped during the Neolithic period.[44]

Ehret cited genetic evidence which had identified the Horn of Africa as a source of a genetic marker “M35/215” Y-chromosome lineage for a significant population component which moved north from that region into Egypt and the Levant. Ehret argued that this genetic distribution paralleled the spread of the Afrasian language family with the movement of people from the Horn of Africa into Egypt and added a new demic component to the existing population of Egypt 17,000 years ago.[45]

See also

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References

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  3. ^ a b c Bender ML (1997). "Upside Down Afrasian". Afrikanistische Arbeitspapiere 50. pp. 19–34.
  4. ^ Militarev A (2005). "Once more about glottochronology and comparative method: the Omotic-Afrasian case, Аспекты компаративистики – 1 (Aspects of comparative linguistics – 1). FS S. Starostin." (PDF). Orientalia et Classica II. Moscow. pp. 339–408. Archived (PDF) from the original on 2020-12-07. Retrieved 2009-07-19.{{cite book}}: CS1 maint: location missing publisher (link)
  5. ^ a b Güldemann, Tom (2018). The Languages and Linguistics of Africa. De Gruyter Mouton. p. 311. ISBN 978-3-11-042606-9.
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  12. ^ Ehret, Christopher; Vyas, Deven N.; Assefa, Shiferaw; Gaston, J. Lafayette; Gleason, Tiffany; Kitchen, Andrew (2023). "Ancient Egyptian's Place in the Afroasiatic Language Family". In Almansa-Villatoro, M. Victoria; Štubňová Nigrelli, Silvia (eds.). Ancient Egyptian and Afroasiatic: Rethinking the Origins. Eisenbrauns. pp. 255–277. ISBN 978-1-64602-212-0.
  13. ^ Scheinfeldt, Laura B.; Soi, Sameer; Tishkoff, Sarah A. (2010-05-11). "Working toward a synthesis of archaeological, linguistic, and genetic data for inferring African population history". Proceedings of the National Academy of Sciences. 107 (supplement_2): 8931–8938. doi:10.1073/pnas.1002563107. ISSN 0027-8424. PMC 3024023. PMID 20445100.
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  15. ^ Militarev, Alexander (2009). "Proto-Afrasian Lexicon Confirming West Asian Homeland: Pastoralism". Journal of Language Relationship. 1: 95–106.
  16. ^ a b Diamond, Jared; Bellwood, Peter (2003-04-25). "Farmers and Their Languages: The First Expansions". Science. 300 (5619): 597–603. Bibcode:2003Sci...300..597D. doi:10.1126/science.1078208. ISSN 0036-8075. PMID 12714734. S2CID 13350469. Archived from the original on 2024-05-26. Retrieved 2023-06-07.
  17. ^ What is Africa? A Human Perspective. (part of: Modern Human Origins and Dispersal, edited by Yonatan Sahle, Hugo Reyes-Centeno, Christian Bentz) April 2019 In book: Modern Human Origins and Dispersal (pp.15–24)Publisher: Kerns Verlag Tübingen https://www.researchgate.net/publication/332735884_What_is_Africa_A_Human_Perspective_part_of_Modern_Human_Origins_and_Dispersal_edited_by_Yonatan_Sahle_Hugo_Reyes-Centeno_Christian_Bentz Archived 2024-05-26 at the Wayback Machine
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  19. ^ Bellwood P, Renfrew C (2002). Examining the farming/language dispersal hypothesis. Cambridge: McDonald Institute for Archaeological Research.
  20. ^ Blench R (2006). Archaeology, Language, and the African Past. AltaMira Press. ISBN 978-0-7591-0466-2.
  21. ^ Diakonoff I (1998). "The Earliest Semitic Society: Linguistic Data". Journal of Semitic Studies. 43 (2): 209–219. doi:10.1093/jss/43.2.209.
  22. ^ Shirai N (2010). The Archaeology of the First Farmer–Herders in Egypt: New Insights into the Fayum Epipalaeolithic and Neolithic. Leiden University Press. ISBN 978-90-8728-079-6.[page needed]
  23. ^ Militarev, Alexander (2009). "Proto-Afrasian Lexicon Confirming West Asian Homeland: Pastoralism". Journal of Language Relationship. 1: 95–106. Archived from the original on 2023-03-23. Retrieved 2023-03-23.
  24. ^ Les langues chamito-sémitiques au Ve siècle avant J.-C. (d’après Meillet et Cohen, 1924).
  25. ^ Campbell, Lyle (2021). Historical Linguistics, Fourth Edition. The MIT Press. pp. 399–400. ISBN 978-0-262-54218-0.
  26. ^ Ehret C (1979). "On the Antiquity of Agriculture in Ethiopia". Journal of African History. 20 (2): 161–177. doi:10.1017/S002185370001700X. JSTOR 181512. S2CID 162986221. Archived from the original on 2022-03-26. Retrieved 2022-03-27 – via JSTOR.
  27. ^ Ehret C (1995). Reconstructing Proto-Afroasiatic (Proto-Afrasian): Vowels, Tone, Consonants, and Vocabulary. University of California Press. ISBN 978-0-520-09799-5.
  28. ^ Ehret C (2002). The Civilizations of Africa: A History to 1800. James Currey Publishers. ISBN 978-0-85255-475-3.
  29. ^ Ehret C (2002). "Language Family Expansions: Broadening our Understandings of Cause from an African Perspective". In Bellwood P, Renfrew C (eds.). Examining the farming/language dispersal hypothesis. Cambridge: McDonald Institute for Archaeological Research.
  30. ^ I. M. Diakonoff (1998) Journal of Semitic Studies 43:209: "It is quite evident that cultural ties between Proto-Semitic and the African branches of the Afrasian macrofamily must have been severed at a very early date indeed. However, the grammatical structure of [Common Semitic] (especially in the verb) is obviously close to that of Common Berbero-Libyan (CBL), as well as to Bedauye. (Bedauye might, quite possibly, be classified as a family distinct from the rest of Kushitic.) The same grammatical isoglosses are somewhat more feebly felt between Semitic and (the other?) Kushitic languages. They practically disappear between the Semitic and the Omotic languages, which were formerly termed Western Kushitic, but which actually may not be Afrasian at all, like their neighbours the Nubian languages and Meroitic."
  31. ^ Rolf Theil (2006) Is Omotic Afro-Asiatic? pp 1–2: "I claim to show that no convincing arguments have been presented [for the inclusion of Omotic (OM) in Afro-Asiatic (AA)], and that OM should be regarded as an independent language family. No closer genetic relations have been demonstrated between OM and AA than between OM and any other language family."
  32. ^ a b Bernal M (1987). Black Athena: the Afroasiatic roots of classical civilization. Rutgers University Press. ISBN 978-0-8135-3655-2.
  33. ^ Bynon, James (1984-01-01). Current Progress in Afro-Asiatic Linguistics: Papers of the Third International Hamito-Semitic Congress. John Benjamins Publishing. pp. 70–73. ISBN 978-90-272-3520-6.
  34. ^ Hodgson, Jason A.; Mulligan, Connie J.; Al-Meeri, Ali; Raaum, Ryan L. (2014-06-12). "Early Back-to-Africa Migration into the Horn of Africa". PLOS Genetics. 10 (6): e1004393. doi:10.1371/journal.pgen.1004393. ISSN 1553-7404. PMC 4055572. PMID 24921250. We hypothesize that a population with substantial Ethio-Somali ancestry could be the proto-Afro-Asiatic speakers. A later migration of a subset of this population back to the Levant before 6 ka would account for a Levantine origin of the Semitic languages [18] and the relatively even distribution of around 7% Ethio-Somali ancestry in all sampled Levantine populations (Table S6). Later migration from Arabia into the HOA beginning around 3 ka would explain the origin of the Ethiosemitic languages at this time [18], the presence of greater Arabian and Eurasian ancestry in the Semitic speaking populations of the HOA (Table 2, S6), and ROLLOFF/ALDER estimates of admixture in HOA populations between 1–5 ka (Table 1). The Ethio-Somali ancestry is found in all admixed HOA ethnic groups, shows little inter-individual variance within these ethnic groups, is estimated to have diverged from all other non-African ancestries by at least 23 ka, and does not carry the unique Arabian lactase persistence allele that arose about 4 ka. Taking into account published mitochondrial, Y chromosome, paleoclimate, and archaeological data, we find that the time of the Ethio-Somali back-to-Africa migration is most likely pre-agricultural.
  35. ^ Vicente, Mário; Schlebusch, Carina M (2020-06-01). "African population history: an ancient DNA perspective". Current Opinion in Genetics & Development. Genetics of Human Origin. 62: 8–15. doi:10.1016/j.gde.2020.05.008. ISSN 0959-437X. PMID 32563853. S2CID 219974966.
  36. ^ Jason A. Hodgson; Connie J. Mulligan; Ali Al-Meeri; Ryan L. Raaum (12 June 2014). "Early Back-to-Africa Migration into the Horn of Africa". PLOS Genetics. 10 (6): e1004393. doi:10.1371/journal.pgen.1004393. PMC 4055572. PMID 24921250.; Supplementary Text S1: Affinities of the Ethio-Somali ancestry component doi:10.1371/journal.pgen.1004393.s017Free access icon
  37. ^ Hodgson JA, Mulligan CJ, Al-Meeri A, Raaum RL (June 2014). "Early back-to-Africa migration into the Horn of Africa". PLOS Genetics. 10 (6): e1004393. doi:10.1371/journal.pgen.1004393. PMC 4055572. PMID 24921250.
  38. ^ McCall, Daniel (February 1998). "The Afroasiatic Language Phylum: African in Origin, or Asian?". Current Anthropology. 39 (1): 139–144. doi:10.1086/204702. ISSN 0011-3204. Archived from the original on 2023-06-01. Retrieved 2022-06-14. My prediction is that Africa will turn out to be the cradle of Afroasiatic, though the speakers of Proto-Afroasiatic were a reflux population from Southwest Asia.
  39. ^ Fregel, Rosa (2021-11-17), "Paleogenomics of the Neolithic Transition in North Africa", Africa, the Cradle of Human Diversity, Brill, pp. 213–235, ISBN 978-90-04-50022-8, archived from the original on 2023-04-18, retrieved 2023-06-07Quote: First, present-day ancestry in North Africans is characterized by an autochthonous Maghrebi component related to a Paleolithic back migration to Africa from Eurasia. ... This result suggests that Iberomaurusian populations in North Africa were related to Paleolithic people in the Levant, but also that migrations of sub-Saharan African origin reached the Maghreb during the Pleistocene.
  40. ^ Baker, Jennifer L.; Rotimi, Charles N.; Shriner, Daniel (2017-05-08). "Human ancestry correlates with language and reveals that race is not an objective genomic classifier". Scientific Reports. 7 (1): 1572. doi:10.1038/s41598-017-01837-7. ISSN 2045-2322. PMC 5431528. PMID 28484253. Arabian ancestry correlates with the Semitic branch of the Afroasiatic language family (r = 0.774, p = 7.28 × 10−51). The Cushitic branch of the Afroasiatic language family correlates with both Eastern African (r = 0.417, p = 7.17 × 10−12) and Arabian (r = 0.336, p = 5.46 × 10−8) ancestries. This result is consistent with our previous finding that Cushitic ancestry formed by admixture between Nilo-Saharan and Arabian ancestries39. ... Northern African ancestry correlates with the Berber branch of the Afroasiatic language family (r = 0.946, p = 1.48 × 10−122). Arabian and Northern African ancestries are both descended from the lineage that includes all Out of Africa migrants, ... Omotic ancestry correlates with the Omotic languages (r = 0.777, p = 1.40 × 10−51). Thus, the genomic data support the linguistic hypothesis that the Omotic languages are not part of the Afroasiatic family42.
  41. ^ Skoglund, Pontus; Thompson, Jessica C.; Prendergast, Mary E.; Mittnik, Alissa; Sirak, Kendra; Hajdinjak, Mateja; Salie, Tasneem; Rohland, Nadin; Mallick, Swapan; Peltzer, Alexander; Heinze, Anja; Olalde, Iñigo; Ferry, Matthew; Harney, Eadaoin; Michel, Megan (September 2017). "Reconstructing Prehistoric African Population Structure". Cell. 171 (1): 59–71.e21. doi:10.1016/j.cell.2017.08.049. ISSN 0092-8674. PMC 5679310. PMID 28938123. While these findings show that a Levant-Neolithic-related population made a critical contribution to the ancestry of present-day eastern Africans (Lazaridis et al., 2016), present-day Cushitic speakers such as the Somali cannot be fit simply as having Tanzania_Luxmanda_3100BP ancestry. The best fitting model for the Somali includes Tanzania_Luxmanda_3100BP ancestry, Dinka-related ancestry, and 16% ± 3% Iranian-Neolithic-related ancestry (p = 0.015). This suggests that ancestry related to the Iranian Neolithic appeared in eastern Africa after earlier gene flow related to Levant Neolithic populations, a scenario that is made more plausible by the genetic evidence of admixture of Iranian-Neolithic-related ancestry throughout the Levant by the time of the Bronze Age (Lazaridis et al., 2016) and in ancient Egypt by the Iron Age (Schuenemann et al., 2017). ... However, this lineage appears to have contributed little ancestry to present-day Bantu speakers in eastern Africa, who instead trace their ancestry to a lineage related to present-day western Africans, with additional components related to the Nilotic-speaking Dinka and to the Tanzania_Luxmanda_3100BP pastoralist (see below; Figure 2).
  42. ^ Prendergast, Mary E.; Lipson, Mark; Sawchuk, Elizabeth A.; Olalde, Iñigo; Ogola, Christine A.; Rohland, Nadin; Sirak, Kendra A.; Adamski, Nicole; Bernardos, Rebecca; Broomandkhoshbacht, Nasreen; Callan, Kimberly; Culleton, Brendan J.; Eccles, Laurie; Harper, Thomas K.; Lawson, Ann Marie (2019-07-05). "Ancient DNA reveals a multistep spread of the first herders into sub-Saharan Africa". Science. 365 (6448). doi:10.1126/science.aaw6275. ISSN 0036-8075. PMC 6827346. PMID 31147405.
  43. ^ Keita, S.O.Y. (ed Bengston, John) (3 December 2008). "Geography, selected Afro-Asiatic families, and Y Chromosome lineage variation: An exploration in linguistics and phylogeography" in In Hot Pursuit of Language in Prehistory: Essays in the four fields of anthropology. In honor of Harold Crane Fleming. John Benjamins Publishing. pp. 3–15. ISBN 978-90-272-8985-8. Archived from the original on 26 May 2024. Retrieved 2 August 2023.{{cite book}}: CS1 maint: multiple names: authors list (link)
  44. ^ Fregel, Rosa (2021-11-17), "Paleogenomics of the Neolithic Transition in North Africa", Africa, the Cradle of Human Diversity, Brill, pp. 213–235, ISBN 978-90-04-50022-8, archived from the original on 2023-04-18, retrieved 2023-08-02 Quote: "Regarding the Y chromosome, current North Africans are characterized by the high frequency of haplogroup E-M81, a lineage considered to be autochthonous of this region. Interestingly, the frequency of E-M81 follows an east to west cline, with the highest frequencies in Morocco and the lowest in Egypt, similar to the results obtained for classical markers. Based on that evidence and contrary to the conclusions drawn from mitochondrial DNA, Arredi et al. (2004) proposed that North African paternal diversity was compatible with a demic expansion from the Middle East. Because the age of E-M81 and other common lineages in North Africa (E-M78 and J-304) were relatively recent, they proposed that the North African pattern of Y-chromosome variation was mostly of Neolithic origin."
  45. ^ Ehret, Christopher (20 June 2023). Ancient Africa: A Global History, to 300 CE. Princeton University Press. pp. 97, 167. ISBN 978-0-691-24410-5. Archived from the original on 15 July 2023. Retrieved 1 August 2023.

Bibliography

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Further reading

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