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Journal of Micropalaeontology

Early Carboniferous (Early Viséan − Serpukhovian) Palynomorphs

G. CLAYTON and S. LOBOZIAK

Journal of Micropalaeontology 1985, v.4; p83-91.

doi: 10.1144/jm.4.1.83

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© The Geological Society of London 2014

 4(1): 83-92,
J.rnicropalaeontol., Downloaded fromMarch 1985
http://jm.lyellcollection.org/ at Memorial University of Newfoundland on August 3, 2014

Early Carboniferous
(Early VisCan - Serpukhovian) Palynomorphs

G. CLAYTON' & S. LOBOZIAK3

'Department of Geology, Trinity College, Dublin, Ireland.
'Laboratoire de Palkobotanique, Sciences de la Terre, Universite des Sciences et Techniques de Lille,
59655 Villeneuve d'Ascq, Cedex, France.

Well preserved assemblages of Early Carboniferous The third and youngest association recorded in Well
miospores have been recovered from a number of wells A1-NC92 in the interval 8115 to 8920ft. and in Well
including A1-NC92, A1-37, A 1 A-84, B1-2, G1-82, A1-14 in the interval 7910 t o 8400ft., is characterised
B1-14 and J1-81A. In some cases the palynomorph by the appearance of monosaccate pollen. In the
assemblages are quantitatively sparse and frequently Rhadames Basin the latter were identified for the first
show evidence of reworking of Late Devonian forms. time in the Eostaffelina and Eosigmolina Biozone
A series of readily indentifiable palynological events (Massa & Vachard, 1979) which is equivalent to the
have been established from examination of Wells J1- Protvian which in turn corresponds with the early part of
81A, A1-NC92, A1-14, A1-37 and A 1 A-84. The the Late Serpukhovian. In Australia, Kemp et al., 1977
oldest miospore assemblage has been recognised in Well have found these monosaccates appearing for the first
A1-37 in the interval 7914 to 8185ft. and in the lowest time at the base of the Anabaculites yberti Assemblage
part of Well A1-NC92 above 9623ft. and is character- Zone, which has been dated independently by cono-
ised by abundant Spelaeotriletes balteatus occurring donts and ostracods (Jones et al., 1973) as Early
together with S . owensi, Radiizonates genuinus and Namurian and by its occurrence with the Echinochonchus
Vallatisporitesagadesi. The assemblage is supplemented gradatus Brachiopod fauna of Roberts (1971) as Late
in the lower part of the interval by Spelaeotriletes VisCan o r Early Namurian.
pretiosus and Vallatisporites vallatus. This association is The interval characterised by this assemblage is
considered to be no older than Early VisCan. Assemblages therefore attributed t o the Serpukhovian.
containing Spelaeotriletes pretiosus and Vallatisporites
vallatus from the Rhadames Basin in western Libya REFERENCES
(Massa et al., 1980) were considered to be Toumaisian Clayton, G., Coquel, R., Doubinger, J., Gueinn, K.J.,
age but the abundance of Spelaeotriletes balteatus in the Loboziak, S., Owens, B. & Streel, M. 1977. Carboniferous
miospores of Western Europe : illustration and zonation.
present samples strongly suggests an earliest VisCan age
Meded. Rijks. Geol. Dienst., 29, 1-71.
to be more probable. Tournaisian sediments probably Jones, P.J.. Campbell, K.S.W. &Roberts, J. 1973. Correlation
occur in some of the wells but the poor state of preser- chart for the Carboniferous System of Australia. Bur. Min.
vation together with problems of reworking d o not at the Res. Australia, Bull., 156A, 1-40.
present time permit them to be positively identified Kemp, E. M., Balme, B.E., Helby, R. J., Kyle, R. A., Playford,
pal ynologicall y. G. & Price, P.L. 1977. Carboniferous and Permian
The second association which is recognised in palynostratigraphy in Australia and Antarctica: a review.
Journ. Austr. Geol. Geophys., 2, 177-208.
Well A1-NC 92 in the interval 8900 t o 9623ft., in Well Massa, D., Coquel, R., Loboziak, S. & Taugourdeau-Lantz, J.
J1-81A and in the lowest part of Well A1-14 is 1980. Essai de synthkse stratigraphique et palynologique du
distinguished by the appearance of representatives of Carbonifkre en Libye occidentale. Ann. SOC. Gtol. Nord.,
the genus Lycospora and Spelaeotriletes triangulus. The 99.429-442.
appearance of these forms more or less coincides with Massa, D. & Vachard, D. 1979. Le Carbonifkre de Libye
the tops of the ranges of Radiizonates genuinus, occidentale. Biostratigraphie et micropalkontologie.
Spelaeotriletes balteatus, S . owensi and Vallatisporites Position dans le domaine tkthysien d'Afrique d u Nord. Rev.
Inst. Fr. Pttrole., 31, 287-333.
agadesi. Broadly similar distribution patterns were Roberts, J. 1971. Devonian and Carboniferous brachiopods
observed in the Rhadames Basin which were dated as from the Bonaparte Gulf Basin, Northwestern Australia.
Late VisCan (Collenia Unit = V3c) and the Early Bur. Min. Res. Australia Bull., 122, 1-319.
Serpukhovian. Representatives of Foveosporites
appositus, Tricidarisporites serratus and Waltzispora
planiangulata are frequently accessory components in
this assemblage.

83
 Downloaded from http://jm.lyellcollection.org/ at Memorial University of Newfoundland on August 3, 2014Clayton & Loboziak

Explanation of Plate 33

All figures are x 500

Fig. 1. Punctutisporites cf. glubrimurginutus Owens, 1971. A1A-84, 5180ft., Slide 1, L31/1, AGC 789. (Frasnian-
VisCan).
Fig. 2. Punctutisporites planus Hacquebard, 1957. A1A-84, 5180ft., Slide 1, P39/1, AGC 790.
Fig. 3. Punctutisporites irrusus Hacquebard, 1957. AlA-84, 5060ft., Slide 1, U30/1, AGC 791.
Fig. 4. Retusotriletes crussus Clayton in Clayton et ul., 1980. A1A-84, 5780ft., Slide 1, Y37/1, AGC 792. (Early
Carboniferous).
Fig. 5. Anapiculutisporites tersus Playford, 1964. J1-81A, 9440-9470ft., Slide 1, U39/1, AGC 793.
Figs. 6, 7. Waltzisporu politu (Hoffmeister, Staplin & Malloy) Smith & Butterworth, 1967. Fig. 6, J1-81A, 8570-
8610ft., Slide 1, W37/3, A G C 794. Fig. 7, J1-81A, 8570-8610ft., Slide 1, Z39/4, AGC 795.
Fig. 8. Umbonatisporites cf. distinctus Clayton, 1970. A1-37, 8185ft., Core 13, S.G. 8185/25, AGC 796.
Fig. 9. Apiculiretusisporu multisetu (Luber) Butterworth & Spinner, 1967. A1-37, 8162ft., Core 13, S.G. 8162/88,
AGC 797. (Early Carboniferous).
Fig. 10. Grunulatisporites grunulutus Ibrahim, 1933. J1-81A, 8570ft., Slide 1, 233, AGC 798.
Fig. 11. Anupiculutisporites concinnus Playford, 1962. 51-81 A, 9140-9180ft., Slide 1 , P37, AGC 799.
Fig. 12. Tricidurisporites dumosus (Staplin) Sullivan & Marshall, 1966. B1-2, 12200-300ft., Slide 1, E32/0,
AGC 800. (VisCan-Namurian).
Fig. 13. Convolutisporu sp. A. A1-37, 8150ft., Core 13, S.G. 8150/32, AGC 801.
Fig. 14. Pustulatisporites sp. A. AlA-84, 5520ft., Slide 1, W37/1, AGC 802.
Fig. 15. Foveosporites uppositus Playford, 1971. 51-81 A, 9140-9180ft., Slide 1, V33/3, AGC 803.
Fig. 16. Pustulatisporites cf. gibberosus Hacquebard, 1957. A1-37, 8162ft., Core 13, S.G. 8162/13, AGC 804.
Fig. 17. Verrucosisporites nitidus (Naumova) Playford, 1963. A1-37, 8145ft., Core 13, Slide 1, P40/2, AGC 805.
(Famennian-VisCan).
Fig. 18. Convolutisporastigmoidea Bharadwaj & Venkatachala, 1961. AlA-84,5340ft., Slide 1, S 5 5 / 0 , AGC 806.
Fig. 19. Convolutispora cf. mellita Hoffmeister, Staplin & Malloy, 1955. A1-37, 8131-32ft., Core 13, S.G. 8131-
8132/56, AGC 807.
Fig. 20. Convolutisporu circumvuflata Clayton, 1970. A1-37, 8162ft., Core 13, Slide 1, P41/1, A G C 808.
Fig. 21. Dictyotriletes sp. A. A1A-84, 5180ft., Slide 1, U38/0, AGC 809.

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G. Clayton & S. Loboziak Plate 33

Early Carboniferous Miospores

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 Downloaded from http://jm.lyellcollection.org/ at Memorial University of Newfoundland on August 3, 2014Clayton & Loboziak

Explanation of Plate 34
All figures are x 500
Figs. 1, 2. Diatornozonotrifetesfragifis Clayton in Neveset a f . , 1973. Fig. 1. AlA-84, 5410ft., Slide 1, W40/2, AGC
810, 2. A1A-84, 5410ft., Slide 1, N32/0, A G C 811.
Fig. 3. Diatomozonotriles sp. A. A1-37, 7960ft., Core 12. S.G. 7960/13, AGC 812.
Fig. 4. Rotaspora knoxi Butterworth & Williams, 1958. J1-81A, 10020-10070ft., Slide 17005, V51/1, AGC 813.
(VisCan-Namurian).
Fig. 5. Cyrtospora cristifera (Luber) Van Der Zwan, 1979. B1-2, 12510-12610ft., Slide 1, Q36/4, AGC 814.
Fig. 6. Densosporites spitsbergensis Playford, 1963. A1A-84, 5340ft., Slide 1, Q62/4, AGC 815.
Fig. 7. Densosporites cf. spinifer Hoffmeister, S t a p h & Malloy, 1955. A1-37, 8162ft., Slide 1, J35/0, AGC 816.
Figs. 8, 14. Densosporites variomarginatus Playford, 1963. Fig. 8, A1-37,8421 ft., S.G. 8421/13, AGC 817. Fig. 14,
A1A-84, 5520ft., Slide 1, Q48/3, AGC 818.
Fig. 9. Densosporites sp. A. A1-37, 8162ft., Core 13, S.G. 8162/9, AGC 819.
Fig. 10. Lycospora pusiffa(Ibrahim) Somers, 1972. A1-NC92, 9500-9600ft., Slide 1, N39/3, AGC 820. (VisCan-
Permian).
Fig. 11.Prolycosporarugufosa(Butterworth &Spinner)Turnau, 1978. A1-14,8790-8900ft., Slide 1, Y52/4, AGC821.
Figs. 12 & 13. Cristatisporites sp. A. Fig. 12. A1-37, 8162ft., Core 13, S.G. 8162/16, AGC 822. Fig. 13. A1-37,
8162ft., Core 13, S.G. 8162/1, AGC 823.
Fig. 15. Densosporites intermedius Butterworth & Williams, 1958. A1-37, 8131-8132ft., Core 13, S.G. 8131-81321
46, AGC 824.
Fig. 16. Cristatisporites sp. B. A1-37, 8162ft., Core 13, S.G. 8162/22, AGC 825.
Figs. 17,22. Spefaeotriletestriangufus Neves & Owens, 1966. Fig. 17. B1-2, 12200-12300ft., Slide 1, H47/1. AGC
826. Fig. 22. J1-81A. 10020-10070ft., Slide 1, N43, AGC 827. (VisCan-Namurian).
Fig. 18. Spefaeotriletes owensi Loboziak & Alpern, 1978. A1A-84, 5180ft., Slide 1, V25/0, AGC 828. (Early
Carboniferous).
Figs.l9,20. Spefaeotriletesbafteatus (Playford) Higgs, 1976. sensu Massa et a f ., 1980. Fig. 19, A1-37,8145ft., Core
13, Slide 1, P41/0, AGC 829. Fig. 20, B1-2, 12200-12300ft., Slide 1, Y27/2, A G C 830. (Early Carboniferous).
Fig. 21. Spelaeotriletes arenaceus Neves & Owens, 1966. AlA-84,568Oft., Slide 1, R44/0, AGC 831. (Tournaisian-
Westphalian).
Fig. 23. Spelaeotriletesprefiosus (Playford) Neves & Belt, 1970. A1-37, 8162ft., Core 13, S.G. 8162/7, AGC 832.

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Early Carboniferous Palynornorphs

G . Clayton & S. Loboziak Plate 34

Early Carboniferous Miospores

87
 Downloaded from http://jm.lyellcollection.org/ at Memorial University of Newfoundland on August 3, 2014 Clayton & Loboziak

Explanation of Plate 35
All figures are x 500
Fig. 1. Spelaeotriletes sp. A. A1-37, 8131-8132ft., Core 13, S.G. 8131-8132165, A G C 833.
Fig. 2. Cirratriradites rarus (Ibrahim) Schopf, Wilson & Bentall, 1944. J1-81A, 9440-9470ft., Slide 1, N4612, A G C
834.
Fig. 3. Spinozonotriletes uncatus Hacquebard, 1957. A1-37, 8162ft., Core 13, S.G. 8162110, AGC 835.
Fig. 4. Radiizonatesgenuinus (Jushko) Loboziak & Alpern, 1978. B1-2,12200-12300ft., Slide 1 , M34/2, A G C 836.
Figs. 5 , 6 . VallatisporitesagadesiLoboziak & Alpern, 1978. Fig. 5. AlA-84,5180ft., Slide 1, U43/2,AGC837. Fig. 6,
A1-37. 8150ft., Core 3, S.G. 815Oll0, A G C 838. (Early Carboniferous).
Figs. 7, 11. Vallatisporites verrucosus Hacquebard, 1957. Fig. 7, A1-37. 8162ft., Core 13, S.G. 8162134, AGC 839.
Fig. 11, B1-2, 12200-12300ft., Slide 1, U44/0, A G C 840.
Fig. 8. Kraeuselisporites echinatus Owens, Mishell & Marshall, 1976. A1-37, 8421ft., S.G. 8421130, AGC 841.
Fig. 9. Vallatisporites vallatus Hacquebard, 1957. A1-37.8162 ft., Core 13, S.G. 8162/27, A G C 842. (Tournaisian-
Early Vistan).
Fig. 10. Vallatisporites cf. ciliaris (Luber) Sullivan, 1964. A1A-84, 5180ft., Slide 1, Q24/0, A G C 843.
Fig. 12. Perotrilites sp. A. A1A-84, 5780ft., Slide 1, T47/0, A G C 844.
Fig. 13. Colatisporitesdecorus (Bharadwaj & Venkatachala) Williams in Nevesetal., 1973. AlA-84,5410ft., Slide 1,
U49/2,AGC 845.
Figs. 14, 15. Spore Type A. Fig. 14, A1-37, 8421 ft., S.G. 8421/29, A G C 846. Fig. 15, A1-37, 8162ft., Core 13,
S.G. 8162/11, AGC 847.

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G . Clayton & S. Loboziak Plate 35

Early Carboniferous Miospores

89
 Downloaded from http://jm.lyellcollection.org/ at Memorial University of Newfoundland on August 3, 2014Clayton & Loboziak

Explanation of Plate 36
All figures are x 500
Fig. 1. Potonieisporites sp. A1-14. 8030-8100ft., Slide 872 (4), S46/2, AGC 783.
Fig. 2. Rimospora rimosa Lele & Maithy, 1969. A1-14, 7910-8000ft., Slide 871 (4), N40/4, AGC 784.
(Serpukhovian-Permian).
Fig. 3. Cannanoropollis janakii PotoniC & Sah, 1958. A1-14, 7910-8000ft., Slide 871 (4), C32/4, AGC 785.
(Serpukhovian- Permian).
Fig. 4. Cannanoropollis janakii PotoniC & Sah, 1958. A1-14, 7910-8000ft., Slide 871 (4), R48/3, AGC 786.
(Serpukhovian -Permian).
Fig. 5 . Plicatipollenites malabarensis (Potonit & Shah, 1958), Foster, 1975. A1-14, 7910-8000ft., Slide 871 (4),
M36/2, A G C 7 87. (Serpukhovian -Permian).
Fig. 6. Barakarites sp. A1-14, 8110-8200ft., Slide 873 (4), G32/3, AGC 788.

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Palynomorphs

S. Loboziak Plate 36
Carboniferous Miospores and Pollen

91