ALEXANDER V.

ZALE
Certified Fisheries Professional

Montana Cooperative Fishery Research Unit, U.S. Geological Survey

Fish & Wildlife Management Program, Department of Ecology
Montana State University, Bozeman, Montana 59717, USA
[email protected]

EDUCATION

University of Florida, Department of Fisheries and Aquaculture

Doctor of Philosophy Degree: Forest Resources and Conservation, December 1984
Dissertation: Applied aspects of the thermal biology, ecology, and life history of the blue
tilapia, Tilapia aurea (Pisces: Cichlidae)

Virginia Polytechnic Institute and State University, Fisheries and Wildlife Sciences
Master of Science Degree: Fisheries and Wildlife, December 1980
Thesis: The life histories of four freshwater lampsiline mussels (Mollusca: Unionidae)
in Big Moccasin Creek, Russell County, Virginia

University of Massachusetts, Department of Forestry and Wildlife Management

Bachelor of Science Degree: Fisheries Biology, June 1978

PROFESSIONAL EXPERIENCE

Montana Cooperative Fishery Research Unit, Biological Resources Division, USGS,

Montana State University, 1/94 - Present

Unit Leader (1/02 - Present) and Professor (12/02 - Present); Assistant Unit Leader
(1/94 - 12/01) and Associate Professor (1/94-12/02): Lead, coordinate, and administer
an applied fisheries research program addressing topics and issues of concern to the
State of Montana and the U.S. Department of the Interior. Responsible for mission
development and execution, proposal preparation, procurement and administration of
research and administrative funds, coordination of joint research efforts with Unit
cooperators, study design, scheduling, analyses, report preparation, supervision of data
acquisition by field personnel, presentations to professional audiences, and publication
of research findings. Technical areas of special emphasis include restoration of native
fishes, whirling disease, thermal ecology, population dynamics and effects of exotic
fishes, and effects of environmental degradation on recreational fisheries.

As Affiliate Professor at Montana State University I teach graduate courses and

seminars in fisheries and aquatic sciences, guest lectures in a variety of graduate and
undergraduate courses, and thesis credits for my advisees. I chair the graduate
committees of my advisees and serve as a member of graduate committees of students
advised by other faculty. In addition to a varying number of graduate students,
postdoctoral associates, and technicians, I supervise the Unit’s Assistant Unit Leader
and Administrative Officer.

Oklahoma Cooperative Fish and Wildlife Research Unit, U.S. Fish and Wildlife
Service, Oklahoma State University, 6/85 - 12/93

Assistant Unit Leader/Fisheries and Adjunct Associate Professor: Coordinated,

supervised, and administered the Oklahoma Unit's applied fisheries research program
addressing topics and issues of concern to the State of Oklahoma and the U.S. Fish
and Wildlife Service. Responsible for proposal preparation, procurement and
administration of research funds, coordination of joint research efforts with Unit
cooperators, study design, scheduling, statistical data analyses, report preparation,
supervision of data acquisition by field personnel, presentations to professional
audiences, and publication of research findings.

Technical areas of special emphasis included effects of hydropower on fisheries, sport

fisheries dynamics in rivers and impoundments of the southern Great Plains and Ozark
Highlands, physiological ecology, early life history, and distribution and ecology of
threatened and endangered fishes. Findings of these investigations were incorporated
by various state and federal agencies into management plans designed to enhance
fisheries resources throughout Oklahoma. Through these investigations, I gained
expertise in regulated-river and reservoir fisheries (turbine and spillway entrainment of
all life history stages, effects of water level fluctuations on reproductive success,
influences of hydropower discharges on reservoir limnology, downstream fish
assemblages and water quality, tailwater recreational fisheries, and reproduction of
riverine fishes), population and exploitation dynamics (creel survey techniques and
analyses, mortality event assessment, effects of harvest regulations, age and growth),
fish assemblage surveys in streams, rivers, and reservoirs, aquatic habitat assessment,
a wide variety of fish collection and enumeration techniques, serological and
hematological field and laboratory techniques, and telemetry and tagging technologies.

As Adjunct Associate Professor at Oklahoma State University I taught one graduate

course entitled Advanced Fisheries Science, special studies courses for graduate
students, guest lectures in a variety of graduate and undergraduate courses, and thesis
hours for my advisees. I chaired 13 graduate committees and served as a member on
an additional 16. My supervisory obligations varied seasonally and as a function of the
number of active projects, but I typically supervised 3 to 5 M.S. candidates, one doctoral
candidate, 5 to 10 part-time temporary undergraduate technicians, and 1 to 3 full-time
technicians. I also supervised one postdoctoral associate and two research associates.

University of Florida, Florida Cooperative Fish and Wildlife Research Unit, 1/85 - 6/85

Postdoctoral Associate: Developed a conceptual framework for the comprehensive

evaluation of existing and potential marsh-management strategies at Merritt Island

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National Wildlife Refuge (Kennedy Space Center), Florida, directed at enhancement of
ecological production and diversity. Compiled and synthesized existing information on
the structure, ecological function, and management of coastal impoundments along the
Atlantic coast of the southeastern United States in the context of their influence on
coastal wetlands habitats and their biota. Supervised the development of an
identification manual for the early life history stages of selected fishes of north-central
Florida, an assessment of the sport fishery of Loxahatchee National Wildlife Refuge,
Florida, and an examination of the potential hazards of diphacinone and
chloraphacinone rodenticides to nontarget fishes. Supervised one biologist, two
technicians, and two MS candidates.

University of Florida, Florida Cooperative Fish and Wildlife Research Unit, 1/84 -
12/84

Biologist: Developed a mathematical Habitat Suitability Index model for use in

evaluation of relative capacities of candidate habitats along the U.S. coast of the Gulf of
Mexico to support nesting by laughing gulls. Supervised laboratory research by a
foreign exchange student on acute effects of environmental pH on blue tilapia.
Prepared and submitted grant proposals and, subsequent to funding, initiated
contracted work prior to official appointment as a postdoctoral associate (see above).

University of Florida, Florida Cooperative Fish and Wildlife Research Unit, 9/80 -
12/83

Graduate Research Assistant: Conducted laboratory and field research on the

distribution, thermal biology, ecology, and life history of the exotic blue tilapia (lower
lethal temperatures, age and growth, reproductive biology, interspecific territoriality,
movements, trophic ecology of early life history stages, electivity by predators).
Supervised two part-time technicians. Assisted in the teaching of undergraduate
courses in fisheries science.

Virginia Polytechnic Institute and State University, Virginia Cooperative Fishery

Research Unit, 9/78 - 9/80

Graduate Research Assistant: Conducted research on the life histories of four

freshwater mussel species in southwestern Virginia (reproductive biology, glochidial
parasitism of host fishes, age and growth). Surveyed distributions of populations of
endangered mussel species in southwestern Virginia. Supervised one full-time
technician.

University of Massachusetts, Massachusetts Cooperative Fishery Research Unit,

3/76 - 9/78 (intermittent appointments)

Fisheries Technician: Assisted U.S. Fish and Wildlife Service personnel and University
of Massachusetts graduate students in research on movements, reproductive biology,
and population ecology of anadromous and endangered fishes of the Connecticut River.

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Crew Chief in 1978. Techniques used: ultrasonic and radio telemetry of adult and
juvenile American shad and Atlantic salmon smolts, ichthyoplankton sampling and
identification, gill netting, electrofishing, beach seining.

GRADUATE STUDENTS SUPERVISED

Completed Theses and Dissertations

Gale, S. B. 2005. Entrainment losses of westslope cutthroat trout into screened and
unscreened irrigation diversion canals on Skalkaho Creek, Montana. M.S. Thesis,
Montana State University, Bozeman. 87 pages.

De Rito, J. N. 2004. Assessment of reproductive isolation between Yellowstone

cutthroat trout and rainbow trout in the Yellowstone River, Montana. M.S. Thesis,
Montana State University, Bozeman. 60 pages.

Lukins, H. J. 2004. Dynamics of the waterborne stage of Myxobolus cerebralis

estimated directly by packed-bed filtration. M.S. Thesis, Montana State University,
Bozeman. 53 pages.

Jaeger, M. E. 2004. An empirical assessment of factors precluding recovery of sauger

in the lower Yellowstone River: movement, habitat use, exploitation, and entrainment.
M.S. Thesis, Montana State University, Bozeman. 115 pages.

Ryce, E. K. N. 2003. Factors affecting the resistance of juvenile rainbow trout to

whirling disease. Ph.D. Dissertation, Montana State University, Bozeman. 124 pages.

Yerk, D. B. 2000. Population characteristics and habitat use of a developing walleye

population in Canyon Ferry Reservoir, Montana. M.S. Thesis, Montana State
University, Bozeman. 195 pages.

Opitz, S. T. 1999. Effects of whirling disease on recruitment of brown trout in the Ruby
River and Poindexter Slough, Montana. M.S. Thesis, Montana State University,
Bozeman. 97 pages.

Mitro, M. G. 1999. Sampling and analysis techniques and their application for
estimating recruitment of juvenile rainbow trout in the Henrys Fork of the Snake River,
Idaho. Ph.D. Dissertation, Montana State University, Bozeman. 288 pages.

Katzman, L. M. 1998. Effects of Predation on Status of Arctic Grayling at Red Rock

Lakes National Wildlife Refuge, Montana. M.S. Thesis, Montana State University,
Bozeman. 207 pages.

Zollweg, E. C. 1998. Piscine predation on bull trout in the Flathead River, Montana.
M.S. Thesis, Montana State University, Bozeman. 97 pages.

4
Simmonds, R. L., Jr. 1995. Effect of piscivorous birds, particularly cormorants, on
reservoir and aquacultural fishes in Oklahoma. M.S. Thesis, Oklahoma State
University, Stillwater. 91 pages.

Shutters, M. K. 1992. Effects of the addition of hydropower to Kaw Dam on striped

bass egg abundances in the Arkansas River, Oklahoma. M.S. Thesis, Oklahoma State
University, Stillwater. 61 pages.

Cunningham, K. K. 1991. Assessment of the effects of environmental variables on the

fishery of the Kaw Lake tailwater, Oklahoma. M.S. Thesis, Oklahoma State University,
Stillwater. 70 pages.

Adornato, T. G. 1990. Comparative influences of environmental parameters on

abundances of hybrid striped bass and white bass in the Grand Lake tailwater,
Oklahoma. M.S. Thesis, Oklahoma State University, Stillwater. 51 pages.

Bohnsack, B. L. 1990. Factors affecting spawning locations and reproductive success

of striped bass in the Arkansas River, Oklahoma. M.S. Thesis, Oklahoma State
University, Stillwater. 83 pages.

Jacks, L. S. 1990. Movements and dynamics of a riverine stock of striped bass

(Morone saxatilis). M.S. Thesis, Oklahoma State University, Stillwater. 47 pages.

Sorenson, K. M. 1990. Entrainment susceptibilities of fishes inhabiting the lower

portion of Grand Lake, Oklahoma. M.S. Thesis, Oklahoma State University, Stillwater.
55 pages.

Stubbs, K. C. 1990. Relative population abundance and size structure dynamics of

white crappie in Skiatook Lake, Oklahoma, during a 5-year stage-fill. M.S. Thesis,
Oklahoma State University, Stillwater. 37 pages.

Travnichek, V. H. 1990. Estimated entrainment of ichthyoplankton through the

Pensacola Dam hydroelectric facility. M.S. Thesis, Oklahoma State University,
Stillwater. 88 pages.

Current Graduate Students

Brown, P. Ph.D. candidate. Evaluation of the efficiency and efficacy of non-native fish
eradication and exclusion techniques for native fish restoration.

Shepard, B. B. Ph.D. candidate. Factors that influence invasion of nonnative brook and
rainbow trout and their interactions with native Yellowstone and westslope cutthroat
trout.

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Davis, W. N. M.S. candidate. Effects of coalbed natural gas development and
extraction on prairie fish assemblages.

Bahn, L. M.S. candidate. Assessment of losses of native fish to irrigation diversions on

selected tributaries of the Bitterroot River, Montana.

Harnish, R. A. M.S. candidate. Efficacy of fish screens and irrigation canal siphons on
Skalkaho Creek, Montana.

Steed, A. C. M.S. candidate. Spatial dynamics of Arctic grayling in the Gibbon River,
Yellowstone National Park.

Spinelli, J. P. M.S. candidate. Factors influencing turbine and spillgate losses of fish
from Hauser Reservoir, Montana.

CONTRACTS AND GRANTS

Evaluation of habitat restoration for the fluvial Arctic grayling in the Big Hole River,
Montana. 2006. Wild Fish Habitat Initiative, Montana University System Water Center
and U.S. Fish and Wildlife Service, $30,430.

Montana prairie riparian native species study: spatiotemporal dynamics of prairie stream
fishes. 2006. Montana Department of Fish, Wildlife & Parks, $101,000.

Effects of coalbed methane development on Great Plains fish assemblages. 2005.

Wild Fish Habitat Initiative, Montana University System Water Center and U.S. Fish and
Wildlife Service, $63,923

Factors influencing turbine and spillgate losses of fish from Hauser Reservoir, Montana.
2005. Montana Department of Fish, Wildlife & Parks and PPL Montana, $90,253

Assessment of losses of native fish to irrigation diversions on selected tributaries of the

Bitterroot River, Montana. Montana Department of Fish, Wildlife & Parks. 2004.
$82,065

Literature review and development of a study plan to assess the effects of coalbed
natural gas activities on fish assemblages. 2004. Bureau of Land Management,
$107,058

Spatial dynamics of Arctic grayling in the Gibbon River, Yellowstone National Park.
2004. National Park Service, $65,752

Movements of resident and non-resident anglers in Montana: implications for

transferring whirling disease among drainages. 2004. National Partnership on the
Management of Wild and Native Cold Water Fisheries, $106,641

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Evaluation of habitat restoration for conservation of cutthroat trout. 2004. Wild Fish
Habitat Initiative, Montana University System Water Center and U.S. Fish and Wildlife
Service, $85,412

Evaluation of the efficiency and efficacy of piscicides for use in non-native fish
eradication. 2003. Wild Fish Habitat Initiative, Montana University System Water
Center and U.S. Fish and Wildlife Service, $115,502

Use of the Powder and Tongue rivers by sauger for spawning. 2003. Montana
Department of Fish, Wildlife & Parks, $25,000

Temporal, spatial, and discharge-mediated dynamics of triactinomyxon abundances and

infection risk estimated directly by packed-bed filtration. 2002. National Partnership on
the Management of Wild and Native Cold Water Fisheries, $55,128

Montana prairie riparian biodiversity survey. 2002. Montana Department of Fish,

Wildlife & Parks, $390,672

Evaluation of entrainment losses of westslope cutthroat trout at private irrigation

diversions on Skalkaho Creek, Montana. 2002. Wild Fish Habitat Initiative, Montana
University System Water Center and U.S. Fish and Wildlife Service, $280,945

Bacterial coldwater disease in westslope cutthroat trout: hatchery epidemiology and

control. 2002. Wild Fish Habitat Initiative, Montana University System Water Center
and U.S. Fish and Wildlife Service, $121,083

Thermal requirements of westslope cutthroat trout. 2002. Wild Fish Habitat Initiative,
Montana University System Water Center and U.S. Fish and Wildlife Service, $83,137

Fish inventories in four park units of the Rocky Mountain Network. 2001. National Park
Service, $20,000

Determination of the maximum weight radio tag that can be implanted in westslope
cutthroat trout without affecting swimming performance: a challenge to the “2% rule.”
2001. Montana University System Water Center, $3,342

Spatial and temporal spawning distributions of Yellowstone cutthroat trout and rainbow
trout in the upper Yellowstone River drainage. 2000. Montana Department of Fish,
Wildlife & Parks, National Fish and Wildlife Foundation, and Jackson Hole One-Fly
Foundation, $88,780

Distribution, movements, and life history characteristics of genetically pure westslope

cutthroat trout Oncorhynchus clarki lewisi in the Fan Creek drainage, Yellowstone
National Park. 2000. U.S. National Park Service and Yellowstone Park Foundation,
$43,750

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Movements and exploitation of sauger in the lower Yellowstone River, Montana. 2000.
Montana Department of Fish, Wildlife & Parks, $120,000

Effects of size versus age on the development of whirling disease in rainbow trout.
2000. National Partnership on the Management of Wild and Native Cold Water
Fisheries, $52,554

Field collecting the triactinomyxon stage of Myxobolus cerebralis using combined

filtration methods. 2000. National Partnership on the Management of Wild and Native
Cold Water Fisheries, $25,878

Comparative use of modified and natural habitats of the upper Yellowstone River by
juvenile salmonids. 1999. U.S. Army Corps of Engineers, $97,536

Assessment of the biological integrity of prairie streams in the Missouri and Yellowstone
River basins, Montana. 1999. U.S. Environmental Protection Agency, $297,813

Fishery survey of the streams of the Charles M. Russell National Wildlife Refuge,
Montana. 1999. Biological Resources Division, U.S. Geological Survey and U.S. Fish
and Wildlife Service, $14,000

The effect of initial exposure to Myxobolus cerebralis on the development of resistance

to reinfection in rainbow trout. 1999. National Partnership on the Management of Wild
and Native Cold Water Fisheries, $61,651

Laboratory assessment of possible selection for resistance to whirling disease among

progeny of Colorado River rainbow trout. 1998. National Partnership on the
Management of Wild and Native Cold Water Fisheries, $22,000

Growth and survival temperature criteria for bull trout. 1998. National Council of the
Paper Industry for Air and Stream Improvement and Biological Resources Division, U.S.
Geological Survey, $81,400

Laboratory investigations of mountain whitefish Prosopium williamsoni susceptibility to

Myxobolus cerebralis. 1998. National Partnership on the Management of Wild and
Native Cold Water Fisheries, $26,870

Effects of age, dose, and environmental stress on development of whirling disease in

rainbow trout. 1997. National Partnership on the Management of Wild and Native Cold
Water Fisheries, $57,568

Development of a method for field collecting the triactinomyxon stage of Myxobolus

cerebralis using paired rotating-drum filtration. 1997. National Partnership on the
Management of Wild and Native Cold Water Fisheries, $22,932

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Effects of whirling disease on recruitment of brown trout in the Ruby River and
Poindexter Slough, Montana. 1996. Montana Department of Fish, Wildlife & Parks,
$57,448

Effects of predation on status of Arctic grayling Thymallus arcticus at Red Rock Lakes
National Wildlife Refuge, Montana. 1995. U.S. National Biological Service and U.S.
Fish and Wildlife Service, $39,000

Development and potential management of an illegally introduced walleye population in

Canyon Ferry Reservoir, Montana. 1995. Montana Department of Fish, Wildlife &
Parks, $40,000

Lake trout predation on juvenile bull trout in the Flathead River System. 1995. Montana
Department of Fish, Wildlife & Parks and Bonneville Power Authority, $17,600

Recruitment of rainbow trout in the Henrys Fork of the Snake River, Idaho. 1994. Idaho
Department of Fish and Game, $108,626

Travel grant to Romania: larval and juvenile fish assemblages as indicators of

environmental health and biological productivity in the Danube Delta. 1994. U.S.
National Academy of Sciences, $2,500

Leopard darter mark and recapture study. 1993. Oklahoma Department of Wildlife
Conservation, $79,998

Travel grant to Latvia: design, development, and initiation of a nationwide ichthyofaunal

survey of Latvian stream and river habitats. 1993. U.S. Fish and Wildlife Service and
Latvian Academy of Sciences, $6,800

Effect of piscivorous birds, particularly cormorants, on reservoir and aquacultural fishes

in Oklahoma. 1992. U.S. Fish and Wildlife Service, $49,590

Travel grant to Latvia: collaborative aquatic environmental assessments. 1992. U.S.

Fish and Wildlife Service and Latvian Academy of Sciences, $6,400

Kaw Hydroelectric Project: Keystone Reservoir recruitment investigations. 1992.

Oklahoma Municipal Power Authority, $4,795

An evaluation of the effect of decreased chloride concentration in the Red River Basin
on primary production and sport fish abundance in Lake Texoma. 1991. U.S. Army
Corps of Engineers, $20,619

Status of the speckled chub in the Arkansas River Basin. 1991. Oklahoma Department
of Wildlife Conservation, $26,666

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Status of the Neosho madtom in Oklahoma. 1991. Oklahoma Department of Wildlife
Conservation, $3,333

Life history and distribution of the Arkansas River shiner in Oklahoma. 1989. Oklahoma
Department of Wildlife Conservation, $60,000

Preliminary studies on the laboratory rearing of leopard darters, Percina pantherina.

1988. U.S. Army Corps of Engineers, $30,000

Career opportunities for minorities in the fisheries and wildlife professions: an

awareness-enhancement workshop for junior high school students. 1988. U.S. Fish
and Wildlife Service, $3,000

Effects of the Pensacola Hydropower Project on the fishery resources of the Grand
River. 1987. Grand River Dam Authority, $403,098

Status of smallmouth bass populations in eastern Oklahoma. 1987. Oklahoma

Department of Wildlife Conservation, $105,191

Kaw Hydroelectric Project: aquatic resources investigations. 1987. Oklahoma

Municipal Power Authority, $182,000

Reproductive ecology of the leopard darter. 1986. Oklahoma Department of Wildlife

Conservation, $80,721

Analysis of the suitability of keystone reservoir, Oklahoma, for habitation by adult striped
bass. 1986. Oklahoma Department of Wildlife Conservation, $57,080

Movements of striped bass in the Arkansas River downstream from Keystone

Reservoir. 1986. Oklahoma Department of Wildlife Conservation, $50,622

Development of a white crappie fishery in a stage-fill impoundment in Oklahoma. 1986.

Oklahoma Department of Wildlife Conservation, $136,132

Movements and harvest of striped bass in the Arkansas River below Keystone Dam.
1986. Oklahoma Department of Wildlife Conservation, $1,725

Telemetric observation of movements of striped bass in the Arkansas River below

Keystone Dam. 1986. Oklahoma Department of Wildlife Conservation, $5,000

Impact of hydropower releases on striped bass in the Arkansas River. 1986. Benham-
Holway Power Group, $1,209

Distribution and abundances of the leopard darter, Percina pantherina (Moore and
Reeves). 1986. U.S. Army Corps of Engineers, $59,000

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Population parameters of striped bass x white bass hybrids in a turbid Oklahoma
reservoir. 1985. Oklahoma Department of Wildlife Conservation, $26,710

Survey of literature on the role of intermittent Great Plains prairie streams in the
dynamics of associated terrestrial and aquatic systems. 1985. U.S. Fish and Wildlife
Service, $4,000

Species profiles of the ladyfish and tarpon, and the reef-building tube worm. 1985.
U.S. Fish and Wildlife Service, $4,000

A conceptual model of salt marsh management on Merritt Island, Florida. 1984.

National Aeronautics and Space Administration, $21,196

Impounded coastal wetlands of the Atlantic coast of the southeastern United States:
literature synthesis and management options. 1984. U.S. Fish and Wildlife Service,
$21,000

PROFESSIONAL AFFILIATIONS

American Fisheries Society

American Society of Ichthyologists and Herpetologists
North American Benthological Society

PROFESSIONAL SERVICE

Lead Co-editor, Fisheries Techniques, 3rd edition, Education Section, American

Fisheries Society, 2005-present
Co-editor, AFS Guide to Fisheries Employment, 2nd edition, Education Section,
American Fisheries Society, 2004-present
Member, Presentation and Poster Guidelines Committee, American Fisheries Society,
2005-present
Member, Meeting Oversight Committee, American Fisheries Society, 2004-present
Member, Award of Excellence Committee, American Fisheries Society, 2002-present
Member, Program Committee, Western Division, American Fisheries Society, 2005-
present
Member (Graphic Designer), Program Committee, North Central Division, American
Fisheries Society, 2002
Associate Editor for Fisheries, Intermountain Journal of Sciences, 1994-2001
Member, Editorial Board, Intermountain Journal of Sciences, 1994-2001
Chair, Nominating Committee, Education Section, American Fisheries Society, 2000
Member, Nominating Committee, American Fisheries Society, 1998-2000
President, Education Section, American Fisheries Society, 1997-1999
Member, Certification Revision Implementation Committee, American Fisheries Society,
1997-1998

11
Member, Special Management Committee, American Fisheries Society, 1997-1998
Member, Program Development Committee, American Fisheries Society, 1997-1998
Member, Certification Revision Task Force, American Fisheries Society, 1996-1997
Chair, Program Committee, Education Section, American Fisheries Society, 1995-1997
Member, Publication Awards Committee, American Fisheries Society, 1995-1996
Chair, Board of Professional Certification, American Fisheries Society, 1994-1996
Secretary-Treasurer, Education Section, American Fisheries Society, 1993-1995
Member, Board of Professional Certification, American Fisheries Society, 1993-1994
Southern Division Representative, Education Section, American Fisheries Society,
1991-1993
Secretary-Treasurer, Introduced Fish Section, American Fisheries Society, 1990-1993
Newsletter Editor, Oklahoma Chapter American Fisheries Society, 1990-1993
Contributed Paper Vice-Chair, 1992 Annual Meeting Program Committee, American
Fisheries Society, 1991-1992
Associate Editor, North American Journal of Fisheries Management, 1990-1992
Member, Warmwater Streams Technical Committee, Southern Division, American
Fisheries Society, 1988-1992
Chair, Purposeful Introductions Committee, Introduced Fish Section, American
Fisheries Society, 1990-1991
Chair, Contributed Paper Session, Program Committee, Southern Division American
Fisheries Society, 1990
President, Oklahoma Chapter American Fisheries Society, 1989-1990
Chair, Student Concerns Committee, Education Section, American Fisheries Society,
1989-1990
Chair, Student Presentation and Poster Awards Committee, Education Section,
American Fisheries Society, 1989-1990
Member, Native Peoples' Fisheries Committee, American Fisheries Society, 1989-1990
Chair, Program Committee, Oklahoma Chapter American Fisheries Society, 1988-1989
Member, Student Presentation and Poster Awards Committee, Education Section,
American Fisheries Society, 1988-1989
Chair, Awards Committee, Oklahoma Chapter American Fisheries Society, 1987-1989
Chair, Membership Committee, Introduced Fish Section, American Fisheries Society,
1987-1988
President, Virginia Tech Chapter American Fisheries Society, 1979-1980

ACADEMIC SERVICE

Faculty Advisor, Fly Fishing Club, Montana State University, 2005-present

Member, Institutional Animal Care and Use Committee, Montana State University,
2004-present
Chief Scientist, Wild Fish Habitat Initiative, Montana Water Center, 2002-present
Co-chair, Department of Ecology Seminar Committee, Montana State University, 2005-
2006
MSU Representative, Aquatic Sciences Laboratory Steering Committee, 1999-present

12
Member, Department of Ecology Executive Committee, Montana State University, 2002-
2003
Chair, A.J.M. Johnson Aquatics Laboratory Design Committee, Montana State
University, 2002

TECHNICAL SERVICE

Member, Montana Cutthroat Technical Committee, 2005-present

Member, Montana Department of Transportation Wildlife and Transportation Technical
Panel, 2000-present
Member, Technical Advisory Committee, Governor’s Upper Yellowstone River Task
Force, 1999-2003
Invited Participant, NPS Greater Yellowstone Network Vital Signs Monitoring Workshop,
2003
Panelist, USEPA Bull Trout Temperature Criteria Peer Review Panel, 2002
Member, Grand Canyon Monitoring and Research Center Review Panel, 1999
Member, Science Sub-committee, Montana Whirling Disease Task Force, 1995-1999
Member, Oklahoma Aquaculture Advisory Group, 1989-1993

HONORS AND AWARDS

2006 Research Grade Promotion, USGS

2005 STAR (Special Thanks for Achieving Results) Award, USGS
2004 STAR Award, USGS
2003 Co-author of Honorable Mention, Best Student Paper at the Annual Meeting of
the American Fisheries Society (Matthew Jaeger)
2003 Leadership Award, National Cooperative Research Units Recognition Awards
Program, USGS
2002 Promoted to Full Professor, Department of Ecology, Montana State University
2001 Co-author of Best Student Paper at the Annual Meeting of the Montana Chapter
of the American Fisheries Society (Eileen Ryce)
2000 Co-author of Outstanding Student Presentation at the Annual Meeting of the
American Fisheries Society (Eileen Ryce)
2000 STAR Award, USGS
1998 Co-author of Honorable Mention, Outstanding Student Presentation at the
Annual Meeting of the American Fisheries Society (Matthew G. Mitro)
1998 Co-author of the Best Student Presentation at the Annual Meeting of the
Montana Chapter American Fisheries Society (Laura Katzman)
1995 Quality Performance Award, U.S. National Biological Service
1995 Research Grade Promotion, U.S. National Biological Service
1994 Quality Performance Award, U.S. National Biological Service

13
1994 Appointed Senior Advisor and Review Panel Committee Member, U.S. National
Academy of Sciences Young Investigator Program on Agricultural Impacts on
Water Quality in Latvia and Lithuania
1993 Special Achievement Award, U.S. Fish and Wildlife Service
1992 Special Achievement Award, U.S. Fish and Wildlife Service
1992 Director's Award for Excellence in Management, presented to the Oklahoma
Cooperative Fish and Wildlife Research Unit, Cooperative Research Units
Center, U.S. Fish and Wildlife Service
1991 Co-author of the Outstanding Fisheries/Wildlife Presentation at the Annual
Meeting of the Oklahoma Academy of Science (William Stark)
1991 Selected for the 1991-92 U.S.--Romanian Summer Program for Young
Investigators in Ecology/Environmental Sciences, U.S. National Academy of
Sciences
1991 Conservation Organization of the Year Award, presented to the Oklahoma
Cooperative Fish and Wildlife Research Unit, Oklahoma Wildlife Federation
1990 Oklahoma Fishery Worker of the Year Award, Oklahoma Chapter American
Fisheries Society
1990 Quality Performance Award, U.S. Fish and Wildlife Service
1989 Co-author of the Outstanding Fisheries Presentation at the Annual Meeting of the
Oklahoma Academy of Science (William Stark)
1989 Special Achievement Award, U.S. Fish and Wildlife Service
1988 Promoted to Associate Professor, Oklahoma State University
1988 Quality Performance Award, U.S. Fish and Wildlife Service
1987 Special Achievement Award, U.S. Fish and Wildlife Service
1978 Graduated Magna Cum Laude, University of Massachusetts, Amherst
1978 Outstanding Fisheries Biology Student Award, University of Massachusetts,
Amherst

PUBLICATIONS

Refereed Journal Articles

Jaeger, M. E., A. V. Zale, T. E. McMahon, and B. J. Schmitz. 2005. Seasonal

movements, habitat use, aggregation, exploitation, and entrainment of sauger in
the lower Yellowstone River: an empirical assessment of factors affecting
population recovery. North American Journal of Fisheries Management 25:1550-
1568.

Zale, A. V., C. Brooke, and W. C. Fraser. 2005. Effects of surgically implanted

transmitter weights on growth and swimming stamina of small adult westslope
cutthroat trout. Transactions of the American Fisheries Society 134:653-660.

14
Bramblett, R. G., T. R. Johnson, A. V. Zale, and D. G. Heggem. 2005. Development
and evaluation of a fish assemblage index of biotic integrity for northwestern
Great Plains streams. Transactions of the American Fisheries Society 134:624-
640.

Ryce, E. K. N., A. V. Zale, E. MacConnell, and M. Nelson. 2005. Effects of fish age
versus size on the development of whirling disease in rainbow trout. Diseases of
Aquatic Organisms 63:69-76.

Ryce, E. K. N., A. V. Zale, and E. MacConnell. 2004. Effects of fish age and parasite
dose on the development of whirling disease in rainbow trout. Diseases of
Aquatic Organisms 59:225-233.

Mitro, M. G., A. V. Zale, and B. A. Rich. 2003. The relation between age-0 rainbow
trout (Oncorhynchus mykiss) abundance and winter discharge in a regulated
river. Canadian Journal of Fisheries and Aquatic Sciences 60:135-139.

Mitro, M. G., and A. V. Zale. 2002. Seasonal survival, movement, and habitat use of
age-0 rainbow trout in the Henrys Fork of the Snake River, Idaho. Transactions
of the American Fisheries Society 131:271-286.

Mitro, M. G., and A. V. Zale. 2002. Estimating abundances of age-0 rainbow trout by
mark-recapture in a medium-sized river. North American Journal of Fisheries
Management 22:188-203.

Selong, J. H., T. E. McMahon, A. V. Zale, and F. T. Barrows. 2001. Effect of

temperature on growth and survival of bull trout, with application of an improved
method for determining thermal tolerance in fishes. Transactions of the
American Fisheries Society 130:1026-1037.

Ryce, E. K. N., A. V. Zale, and R. B. Nehring. 2001. Lack of selection for resistance to
whirling disease among progeny of Colorado River rainbow trout. Journal of
Aquatic Animal Health 13:63-68.

Mitro, M. G., and A. V. Zale. 2000. Use of distance sampling to estimate rainbow trout
redd abundances in the Henry’s Fork of the Snake River, Idaho. Intermountain
Journal of Sciences 6:223-231.

Bramblett, R. G., and A. V. Zale. 2000. The ichthyofauna of small streams on the
Charles M. Russell National Wildlife Refuge, Montana. Intermountain Journal of
Sciences 6:57-67.

Zale, A. V., R. L. Simmonds, Jr., and R. T. Eades. 2000. Getting a job or assistantship:
How to surpass the competition. Fisheries 25(6):24-31.

15
Mitro, M. G., and A. V. Zale. 2000. Predicting fish abundance using single-pass
removal sampling. Canadian Journal of Fisheries and Aquatic Sciences 57:951-
961.

Simmonds, R. L., Jr., A. V. Zale, and D. M. Leslie, Jr. 2000. Modeled effects of double-
crested cormorant predation on simulated reservoir sport and forage fish
populations in Oklahoma. North American Journal of Fisheries Management
20:168-179.

Cunningham, K. K., and A. V. Zale. 1998. Dynamics of white crappie exploitation in an

Oklahoma tailwater fishery. Proceedings of the Oklahoma Academy of Science
78:35-40.

Sorenson, K. M., W. L. Fisher, and A. V. Zale. 1998. Turbine passage of juvenile and
adult fish at a warmwater hydroelectric facility in northeastern Oklahoma:
Monitoring associated with relicensing. North American Journal of Fisheries
Management 18:124-136.

Simmonds, R. L., Jr., A. V. Zale, and D. M. Leslie, Jr. 1997. Physical and biological
factors affecting seasonal double-crested cormorant densities on Oklahoma
reservoirs. Colonial Waterbirds 20:31-40.

Zale, A. V., and T. G. Adornato. 1996. Comparative influences of abiotic variables on

seasonal abundances of hybrid striped bass and white bass in the Grand Lake
tailwater, Oklahoma. Journal of Freshwater Ecology 11:257-269.

Baker, J., and eleven coauthors. 1995. Agricultural impacts on water quality in Latvia
and Lithuania: a report of the U.S. Young Investigator Program. Proceedings of
the Latvian Academy of Sciences, Section B 1995:128-136. Invited.

Bohnsack, B. L., and A. V. Zale. 1994. Spatial and temporal distributions of striped
bass eggs in the Arkansas River, Oklahoma, 1987 and 1988. Proceedings of the
Annual Conference Southeastern Association of Fish and Wildlife Agencies
46:458-468.

Zale, A. V., and M. B. Bain. 1994. Estimating tag-reporting rates with postcards as tag
surrogates. North American Journal of Fisheries Management 14:208-211.

Zale, A. V., S. C. Leon, M. Lechner, O. E. Maughan, M. T. Ferguson, S. O'Donnell, B.

James, and P. W. James. 1994. Distribution of the threatened leopard darter,
Percina pantherina (Osteichthyes: Percidae). The Southwestern Naturalist
39:11-20.

Travnichek, V. H., A. V. Zale, and W. L. Fisher. 1993. Entrainment of ichthyoplankton

by a warmwater hydroelectric facility. Transactions of the American Fisheries
Society 122:709-716.

16
Fisher, W. L., and A. V. Zale. 1993. Effect of water level fluctuations on abundance of
young-of-year largemouth bass in a hydropower reservoir. Proceedings of the
Annual Conference Southeastern Association of Fish and Wildlife Agencies
45:422-431.

Luttrell, G. R., R. D. Larson, W. J. Stark, N. A. Ashbaugh, A. A. Echelle, and A. V. Zale.

1992. Status and distribution of the Neosho madtom (Noturus placidus) in
Oklahoma. Proceedings of the Oklahoma Academy of Science 72:5-6.

James, P. W., O. E. Maughan, and A. V. Zale. 1991. Life history of the leopard darter
Percina pantherina in Glover River, Oklahoma. American Midland Naturalist
125:173-179.

Zale, A. V., J. D. Wiechman, R. L. Lochmiller, and J. Burroughs. 1990. Limnological

conditions associated with summer mortality of striped bass in Keystone
Reservoir, Oklahoma. Transactions of the American Fisheries Society 119:72-
76.

Zale, A. V., and R. W. Gregory. 1990. Food selection by early life stages of blue tilapia,
Oreochromis aureus, in Lake George, Florida: overlap with sympatric shad
larvae. Florida Scientist 53:123-129.

Conrow, R., A. V. Zale, and R. W. Gregory. 1990. Distributions and abundances of

early life stages of fishes in a Florida lake dominated by aquatic macrophytes.
Transactions of the American Fisheries Society 119:521-528.

Zale, A. V., and R. W. Gregory. 1989. Effect of salinity on cold tolerance of juvenile
blue tilapias. Transactions of the American Fisheries Society 118:718-720.

Lochmiller, R. L., J. D. Wiechman, and A. V. Zale. 1989. Hematological assessment of

temperature and oxygen stress in a reservoir population of striped bass (Morone
saxatilis). Comparative Biochemistry and Physiology A: Comparative Physiology
93:535-541.

Zale, A. V. 1987. Growth, survival, and foraging abilities of early life history stages of
blue tilapia, Oreochromis aureus, and largemouth bass, Micropterus salmoides.
Environmental Biology of Fishes 20:113-128.

Zale, A. V. 1987. Periodicity of habitation of a stenothermal spring run in north-central

Florida by blue tilapia. North American Journal of Fisheries Management 7:575-
579.

Montague, C. L., A. V. Zale, and H. F. Percival. 1987. Ecological effects of coastal

marsh impoundments: a review. Environmental Management 11:743-756.

17
James, P., S. Leon, A. V. Zale, and O. E. Maughan. 1987. Diver-operated
electrofishing device. North American Journal of Fisheries Management 7:597-
598.

Schramm, H. L., Jr., and A. V. Zale. 1985. Effects of cover and prey size on
preferences of juvenile largemouth bass for blue tilapias and bluegills in tanks.
Transactions of the American Fisheries Society 114:725-731.

Neves, R. J., L. R. Weaver, and A. V. Zale. 1985. An evaluation of host fish suitability
for glochidia of Villosa vanuxemi and V. nebulosa (Pelecypoda: Unionidae).
American Midland Naturalist 113:13-19.

Zale, A. V., and R. J. Neves. 1982. Fish hosts of four species of lampsiline mussels
(Mollusca: Unionidae) in Big Moccasin Creek, Virginia. Canadian Journal of
Zoology 60:2535-2542.

Zale, A. V., and R. J. Neves. 1982. Reproductive biology of four freshwater mussel
species (Mollusca: Unionidae) in Virginia. Freshwater Invertebrate Biology (now
Journal of the North American Benthological Society) 1:17-28.

Zale, A. V., and R. J. Neves. 1982. Identification of a host fish for Alasmidonta minor
(Mollusca: Unionidae). American Midland Naturalist 107:386-388.

Neves, R. J., and A. V. Zale. 1982. Freshwater mussels (Unionidae) of Big Moccasin
Creek, southwestern Virginia. Nautilus 96:52-54.

Refereed Book Chapters

Zale, A. V. 2006. Pursuing graduate studies in fisheries. Pages 39-55 in D. A. Hewitt,

W. E. Pine, III, and A. V. Zale, editors. 2006. The AFS guide to fisheries
employment, 2nd edition. American Fisheries Society, Bethesda, Maryland.

Katzman, L. M., and A. V. Zale. 2000. Age and growth of an unexploited burbot
population in Upper Red Rock Lake, Montana. Pages 139-146 in V. L.
Paragamian and D. W. Willis, editors. Burbot: biology, ecology, and
management. Fisheries Management Section, American Fisheries Society.
Arthur H. Clark Company, Spokane, Washington. Invited.

McMahon, T. E., A. V. Zale, and D. J. Orth. 1996. Aquatic habitat measurements.

Pages 83-120 in B. R. Murphy and D. W. Willis, editors. Fisheries techniques,
2nd edition. American Fisheries Society, Bethesda, Maryland. Invited.

Echelle, A. A., G. R. Luttrell, R. D. Larson, A. V. Zale, W. L. Fisher, and D. M. Leslie, Jr.

1995. Declines of native prairie fishes. Pages 303-305 in E. T. LaRoe, G. S.
Farris, C. E. Puckett, P. D. Doran, and M. J. Mac, editors. Our living resources:
a report to the nation on the distribution, abundance, and health of U.S. plants,

18
 animals, and ecosystems. U.S. Department of the Interior, National Biological
Service, Washington, DC.

Zale, A. V., O. E. Maughan, D. J. Orth, and W. Layher. 1993. Withdrawals. Pages

271-285. In C. F. Bryan and D. A. Rutherford, editors. Impacts on warmwater
streams: guidelines for evaluation. Southern Division, American Fisheries
Society, Little Rock, Arkansas. Invited.

Refereed Articles in Symposia

Kerans, B. L., and A. V. Zale. 2002. The ecology of Myxobolus cerebralis. Pages 145-
166 in J. L. Bartholomew and J. C. Wilson, editors. Whirling disease: reviews
and current topics. American Fisheries Society, Symposium 29, Bethesda,
Maryland. Invited.

Simmonds, R. L., Jr., A. V. Zale, and D. M. Leslie, Jr. 1995. Depredation of catfish by
double-crested cormorants at aquaculture facilities in Oklahoma. Pages 34-37 in
R. E. Masters and J. G. Huggins, editors. Twelfth Great Plains Wildlife Damage
Control Workshop Proceedings. Great Plains Agricultural Council Publication
153.

Zale, A. V., C. L. Montague, and H. F. Percival. 1987. A synthesis of potential effects

of coastal impoundments on the production of estuarine fish and shellfish and
some management options. Pages 424-436 in W. R. Whitman and W. H.
Meredith, editors. Waterfowl and Wetlands Symposium: Proceedings of a
Symposium on Waterfowl and Wetlands Management in the Coastal Zone of the
Atlantic Flyway. Delaware Coastal Management Program, Delaware Department
of Natural Resources and Environmental Control. Dover, Delaware.

Montague, C. L., A. V. Zale, and H. F. Percival. 1987. The nature of export from
fringing marshes, with reference to the production of estuarine animals and the
effect of impoundments. Pages 437-450 in W. R. Whitman and W. H. Meredith,
editors. Waterfowl and Wetlands Symposium: Proceedings of a Symposium on
Waterfowl and Wetlands Management in the Coastal Zone of the Atlantic Flyway.
Delaware Coastal Management Program, Delaware Department of Natural
Resources and Environmental Control. Dover, Delaware.

Percival, H. F., A. V. Zale, and C. L. Montague. 1987. A summary of positive and

negative aspects of coastal wetland impoundments as habitat for waterfowl.
Pages 223-230 in W. R. Whitman and W. H. Meredith, editors. Waterfowl and
Wetlands Symposium: Proceedings of a Symposium on Waterfowl and Wetlands
Management in the Coastal Zone of the Atlantic Flyway. Delaware Coastal
Management Program, Delaware Department of Natural Resources and
Environmental Control. Dover, Delaware.

19
Zale, A. V., and R. Suttles. 1987. Mussel mortalities in the Neosho River System,
Oklahoma. Pages 39-43 in R. J. Neves, editor. Proceedings of the Workshop on
Die-offs of Freshwater Mussels in the United States. 23-25 June 1986,
Davenport, Iowa.

Editorships

Hewitt, D. A., W. E. Pine, III, and A. V. Zale, editors. 2006. The AFS guide to fisheries
employment, 2nd edition. American Fisheries Society, Bethesda, Maryland.

R. W. Van Kirk, and A. V. Zale. 2000. Aquatic resources of the Henry's Fork
watershed. Intermountain Journal of Sciences 6:103-332 (14 articles).

Zale, A. V. 1998. The relationship between land management activities and habitat
requirements of bull trout. Montana Bull Trout Scientific Group for the Montana
Bull Trout Restoration Team. Montana Department of Fish, Wildlife & Parks,
Helena, Montana. 78 pages.

Zale, A. V., D. L. Parrish, and M. E. Mather. 1998. Enhancing graduate education and
mentoring. Fisheries 23(9):6-17 (4 articles).

Non-Refereed

Davis, W. N., R. G. Bramblett, and A. V. Zale. 2006. Literature review and

development of a study plan to assess the effects of coalbed natural gas
activities on fish assemblages. Final Report to the U.S. Bureau of Land
Management, Miles City, Montana. 79 pages.

Shepard, B. B., and A.. V. Zale. 2005. Dyce Creek brook trout removal project: 2004 to
2005. Final Report to the U.S. Bureau of Land Management, Dillon, Montana.
20 pages.

Gale, S. B., and A. V. Zale. 2005. Evaluation of entrainment losses of westslope

cutthroat trout at private irrigation diversions and the efficiency of fish screens on
Skalkaho Creek, Montana. Final Report to The Wild Fish Habitat Initiative,
Montana University System Water Center, Bozeman, Montana. 75 pages.

Bear, B. A., T. A. McMahon, and A. V. Zale. 2004. Thermal requirements of westslope

cutthroat trout. Final Report to The Wild Fish Habitat Initiative, Montana
University System Water Center, Bozeman, Montana. 31 pages.

Ryce, E. K. N., and A. V. Zale. 2004. Bacterial coldwater disease in westslope

cutthroat trout: hatchery epidemiology and control. Final Report to The Wild
Fish Habitat Initiative, Montana University System Water Center, Bozeman,
Montana. 13 pages.

20
Lukins, H. J., A. V. Zale, and F. T. Barrows. 2004. Triactinomyxon dynamics estimated
by packed-bed filtration. Pages 9-10 in Proceedings of the 10th Annual Whirling
Disease Symposium. Whirling Disease Foundation, Bozeman, Montana.

Bergstedt, L. C., R. G. White, and A. V. Zale. 2004. Development of an index of biotic

integrity for measuring biological condition of the Missouri River. Volume 7.
Population structure and habitat use of benthic fises along the Missouri and lower
Yellowstone rivers. U.S. Geological Survey, Cooperative Research Units. Final
report to the U.S. Army Corps of Engineers, Omaha, Nebraska. 78 pages.

Lukins, H. J., A. V. Zale, and F. T. Barrows. 2003. Temporal, spatial, and discharge
mediated dynamics of of triactinomyxon abundances and infection risk estimated
directly by packed-bed filtration. Final report to the Whirling Disease Initiative of
the National Partnership on the Management of Wild and Native Cold Water
Fisheries, Bozeman, Montana. 26 pages.

Bramblett, R. G., T. R. Johnson, A. V. Zale, and D. Heggem. 2003. Development of

biotic integrity indices for prairie streams in Montana using fish,
macroinvertebrate, and diatom assemblages. Final report to U.S. Environmental
Protection Agency. 96 pages.

Zale, A. V. 2003. Distribution, movements, and life history characteristics of genetically

pure westslope cutthroat trout Oncorhynchus clarki lewisi in the Fan Creek
drainage, Yellowstone National Park. Final report to National Park Service and
Yellowstone Park Foundation. 103 pages.

Lukins, H. J., A. V. Zale, and F. T. Barrows. 2003. Temporal fluctuations in

triactinomyxon densities. Pages 41-42 in Proceedings of the 9th Annual Whirling
Disease Symposium–Managing the Risk. Whirling Disease Foundation,
Bozeman, Montana.

Zale, A. V., and D. Rider. 2003. Comparative use of modified and natural habitats of
the Upper Yellowstone River by juvenile salmonids. Final report to the U.S. Army
Corps of Engineers, Omaha, Nebraska. 66 pages.

Bramblett, R. G., and A. V. Zale. 2002. Fish inventories in four Park Units of the Rocky
Mountain Network. Final Report to the Intermountain Region, National Park
Service, Denver, Colorado. 45 pages.

Zale, A. V., C. Brooke, and W. C. Fraser. 2002. Determination of the maximum-weight

telemetry transmitter that can be implanted in westslope cutthroat trout without
affecting swimming performance: a challenge to the “2% rule.” Final report to
the Montana Water Center, Montana State University, Bozeman, Montana. 16
pages.

21
Leathe, S., G. Liknes, T. McMahon, A. Munro, E. R. Vincent, and A. Zale. 2002.
Epidemiology of whirling disease in the Missouri River, Montana: disease spread
and trout population response five years after discovery. Pages 19-20 in
Proceedings of the 8th Annual Whirling Disease Symposium. Whirling Disease
Foundation, Bozeman, Montana.

Ryce, E. K. N., A. V. Zale, E. MacConnell, and M. A. Adkison. 2002. Factors affecting

the resistance of juvenile rainbow trout to whirling disease. Pages 27-28 in
Proceedings of the 8th Annual Whirling Disease Symposium. Whirling Disease
Foundation, Bozeman, Montana.

Barrows, F. T., A. V. Zale, H. Lukins, and T. E. McMahon. 2002. Validation of packed-

bed filtration for triactinomyxon quantification: laboratory and field trials. Pages
65-66 in Proceedings of the 8th Annual Whirling Disease Symposium. Whirling
Disease Foundation, Bozeman, Montana.

Barrows, F. T., A. V. Zale, H. Lukins, and T. E. McMahon. 2001. Field collecting the
triactinomyxon stage of Myxobolus cerebralis using combined filtration methods.
Final report to the Whirling Disease Initiative of the National Partnership on the
Management of Wild and Native Cold Water Fisheries, Bozeman, Montana. 5
pages.

Ryce, E. K. N., and A. V. Zale. 2001. Effects of size versus age on the development of
whirling disease in rainbow trout. Final report to the Whirling Disease Initiative of
the National Partnership on the Management of Wild and Native Cold Water
Fisheries, Bozeman, Montana. 11 pages.

Leathe, S., G. Liknes, T. McMahon, A. Munro, and A. V. Zale. 2001. Epidemiology of

whirling disease in the Missouri River Ecosystem. Final report to the Whirling
Disease Initiative of the National Partnership on the Management of Wild and
Native Cold Water Fisheries, Bozeman, Montana. 16 pages.

R. W. Van Kirk, and A. V. Zale. 2000. Preface: Aquatic resources of the Henry's Fork
watershed. Intermountain Journal of Sciences 6:103-105.

Craig, A. J., and A. V. Zale. 2001. Effects of bank stabilization structures on fish and
their habitat: a literature review. Report to Omaha District, U.S. Army Corps of
Engineers and Governor’s Upper Yellowstone River Task Force. 27 pages.

Ryce, E. K. N., E. MacConnell, and A. V. Zale. 2001. Effects of size versus age on the
development of whirling disease in rainbow trout. Pages 35-36 in Proceedings of
the 7th Annual Whirling Disease Symposium. Whirling Disease Foundation,
Bozeman, Montana.

Ryce, E. K. N., E. MacConnell, A. V. Zale, and M. A. Adkison. 2001. Effect of initial

exposure to Myxobolus cerebralis on the development of resistance to reinfection

22
 in rainbow trout. Pages 37-38 in Proceedings of the 7th Annual Whirling Disease
Symposium. Whirling Disease Foundation, Bozeman, Montana.

Kerans, B. L., and A. V. Zale. 2001. The ecology of Myxobolus cerebralis. Page 53 in
Proceedings of the 7th Annual Whirling Disease Symposium. Whirling Disease
Foundation, Bozeman, Montana. Invited.

Ryce, E. K. N., A. V. Zale, and E. MacConnell. 2001. The effect of initial exposure to
Myxobolus cerebralis on the development of resistance to reinfection in rainbow
trout. Final report to the Whirling Disease Initiative of the National Partnership on
the Management of Wild and Native Cold Water Fisheries, Bozeman, Montana.
10 pages.

MacConnell, E., A. V. Zale, and M. A. Quinn. 2000. Susceptibility of mountain whitefish

to Myxobolus cerebralis. Pages 107-108 in Proceedings of the 6th Annual
Whirling Disease Symposium. Whirling Disease Foundation, Bozeman,
Montana.

Barrows, F. T., A. V. Zale, and T. E. McMahon. 2000. Triactinomyxon collection using

packed-bed filtration: Update on progress. Pages 81-82 in Proceedings of the
6th Annual Whirling Disease Symposium. Whirling Disease Foundation,
Bozeman, Montana.

Ryce, E. K. N., A. V. Zale, and R. B. Nehring. 2000. Laboratory assessment of

possible selection for resistance to whirling disease among progeny of Colorado
River rainbow trout (Oncorhynchus mykiss). Final report to the Whirling Disease
Initiative of the National Partnership on the Management of Wild and Native Cold
Water Fisheries, Bozeman, Montana. 6 pages.

MacConnell, E., and A. V. Zale. 2000. Susceptibility of mountain whitefish, Prosopium

williamsoni, to Myxobolus cerebralis. Final report to the Whirling Disease
Initiative of the National Partnership on the Management of Wild and Native Cold
Water Fisheries, Bozeman, Montana. 5 pages.

Bramblett, R. G., and A. V. Zale. 2000. Fishery survey of the streams of the Charles M.
Russell National Wildlife Refuge, Montana. Final report to U.S. Geological
Survey. 42 pages.

Ryce, E. K. N., E. MacConnell, and A. V. Zale. 1999. Effects of age, dose and
environmental stress on the development of whirling disease in rainbow trout.
Final report to the Whirling Disease Initiative of the National Partnership on the
Management of Wild and Native Cold Water Fisheries, Bozeman, Montana. 10
pages.

Adelman, I. R., M. L. Armstrong, T. L. Close, W. J. Daley, H. L. Forsgren, P. C.

Jacobson, S. L. McMullin, R. A. Rosen, C. G. Scalet, and A. V. Zale. 1999. The

23
 American Fisheries Society professional certification program. Fisheries
24(9):36-37.

Barrows, F. T., A. V. Zale, T. McMahon, and R. P. Hedrick. 1999. Development of a

field collection method for the triactinomyxon stage of Myxobolus cerebralis.
Pages 19-22 in Proceedings of the 5th Annual Whirling Disease Symposium:
Research & Management Perspectives. Whirling Disease Foundation,
Bozeman, Montana.

Ryce, E. K. N., E. MacConnell, and A. V. Zale. 1999. Effects of age and dose on the
development of whirling disease in rainbow trout. Pages 35-40 in Proceedings of
the 5th Annual Whirling Disease Symposium: Research & Management
Perspectives. Whirling Disease Foundation, Bozeman, Montana.

Ryce, E. K. N., E. MacConnell, and A. V. Zale. 1999. Effects of whirling disease on

rainbow trout endurance and swimming performance. Pages 41-44 in
Proceedings of the 5th Annual Whirling Disease Symposium: Research &
Management Perspectives. Whirling Disease Foundation, Bozeman, Montana.

Ryce, E. K. N., A. V. Zale, and R. B. Nehring. 1999. Selection for resistance to whirling
disease among progeny of Colorado River rainbow trout. Pages 57-50 in
Proceedings of the 5th Annual Whirling Disease Symposium: Research &
Management Perspectives. Whirling Disease Foundation, Bozeman, Montana.

McMahon, T. E., A. R. Munro, D. Downing, E. R. Vincent, S. Leathe, G. Grisak, G.

Liknes, P. Clancey, B. B. Shepard, and A. V. Zale. 1999. Life history variation in
rainbow trout in relation to whirling disease infection risk. Pages 109-111 in
Proceedings of the 5th Annual Whirling Disease Symposium: Research &
Management Perspectives. Whirling Disease Foundation, Bozeman, Montana.

Katzman, L. M., and A. V. Zale. 1998. Effects of predation on status of Arctic grayling
Thymallus arcticus at Red Rock Lakes National Wildlife Refuge, Montana.
Completion Report to Biological Resources Division, U.S. Geological Survey.

Ryce, E. K. N., E. MacConnell, and A. V. Zale. 1998. Effects of age, dose, and
environmental stress on the development of whirling disease in rainbow trout.
Pages 33-36 in Proceedings of the 1998 Whirling Disease Symposium:
Research in Progress. Whirling Disease Foundation, Bozeman, Montana.

Munro, A. R., T. E. McMahon, B. L. Kerans, A. V. Zale, S. Leathe, and E. R. Vincent.

1998. Relation of life history type to whirling disease susceptibility in Missouri
River rainbow trout: Plans and progress. Pages 37-39 in Proceedings of the
1998 Whirling Disease Symposium: Research in Progress. Whirling Disease
Foundation, Bozeman, Montana.

24
Opitz, S. T., and A. V. Zale. 1998. Brown trout recruitment and whirling disease in the
Ruby River and Poindexter Slough, Montana. Pages 59-71 in Proceedings of
the 1998 Whirling Disease Symposium: Research in Progress. Whirling Disease
Foundation, Bozeman, Montana.

Adelman, I. R., M. L. Armstrong, T. L. Close, W. J. Daley, H. L. Forsgren, P. C.

Jacobson, S. L. McMullin, R. A. Rosen, C. G. Scalet, and A. V. Zale. 1997. The
revised American Fisheries Society professional certification program. Fisheries
22(8):11-15.

Opitz, S. T., and A. V. Zale. 1997. Recruitment of brown trout in the Ruby River and
Poindexter Slough, Montana: 1996 data. Pages 79-81 in Whirling Disease
Symposium--Expanding the Database: 1996 Research Progress Reports.
Whirling Disease Foundation, Bozeman, Montana.

Simmonds, R. L., Jr., A. V. Zale, and D. M. Leslie, Jr. 1995. Effect of piscivorous birds,
particularly cormorants, on reservoir and aquacultural fishes in Oklahoma. Final
Report to U.S. Fish and Wildlife Service, Region 2, Albuquerque, New Mexico.
91 pages.

Baker, J., and eleven coauthors. 1995. Young Investigator Program on agricultural
impacts on water quality in Latvia and Lithuania. 1994 Summary Report.
National Academy Press, Washington, DC. 73 pages.

Luttrell, G. R., A. A. Echelle, and A. V. Zale. 1993. Status of the speckled chub in the
Arkansas River Basin. Final Report, Federal Aid Project E-8 (Oklahoma), Job 3.
37 pages.

Zale, A. V. 1993. Keystone Reservoir recruitment investigations. Final Report to

Oklahoma Municipal Power Authority, Edmond, Oklahoma. 10 pages.

Zale, A. V. 1992. Latvian--U.S. collaborative hydrobiological research in Latgale and

Kurzeme. Proceedings of the Latvian Academy of Sciences 1992:76.

Zale, A. V., K. Sorenson, and J. Burroughs. 1992. Analysis of the suitability of

Keystone Reservoir, Oklahoma, for habitation by adult striped bass. Final
Report, Federal Aid Project F-41-R (Oklahoma), Job XII. 55 pages.

Zale, A. V., M. K. Shutters, B. L. Bohnsack, and K. K. Cunningham. 1992. Kaw

Hydroelectric Project aquatic resources investigations: striped bass
investigation. Final Report to Oklahoma Municipal Power Authority, Edmond,
Oklahoma. 120 pages.

Gade, D., S. L. Burks, and A. V. Zale. 1992. An evaluation of the effect of decreased
chloride concentration in the Red River Basin on primary production and sport

25
 fish abundance in Lake Texoma. Final Report to Tulsa District, U.S. Army Corps
of Engineers, Tulsa, Oklahoma. 21 pages.

Zale, A. V., and K. C. Stubbs. 1991. Development of a white crappie fishery in a stage-
fill impoundment in Oklahoma. Final Report, Federal Aid Project F-41-R
(Oklahoma), Job XIII. 81 pages.

Larson, R. D., A. A. Echelle, and A. V. Zale. 1991. Life history and distribution of the
Arkansas River shiner in Oklahoma. Final Report, Federal Aid Project E-8
(Oklahoma), Job 1. 36 pages.

Luttrell, G. R., A. A. Echelle, and A. V. Zale. 1991. Status of the Neosho madtom in
Oklahoma. Final Report, Federal Aid Project E-8 (Oklahoma), Job 2. 22 pages.

Pringle, C., B. Harvey, and A. Zale. 1991. The imperiled Danube delta--U.S.-Romanian
environmental reconnaissance of Europe's largest wetland. Bulletin of the North
American Benthological Society 8:329-335.

Griffin, C., and nine coauthors. 1991. Scientific report of the U.S.-Romanian summer
program for young investigators in ecology/environmental sciences. Final Report
to U.S. National Academy of Sciences, Washington, D.C. 34 pages.

Stark, W. J., and A. V. Zale. 1991. Status of smallmouth bass populations in eastern
Oklahoma. Final Report, Federal Aid Project F-41-R (Oklahoma), Job XIV. 42
pages.

Stark, W. J., A. V. Zale., and D. M. Leslie, Jr. 1991. Unique electrofishing boat for
remote streams. U.S. Fish & Wildlife Service Research Information Bulletin 91-
10.

Berkhouse, C. S., T. M. Brandt, and A. V. Zale. 1990. Preliminary studies on the

laboratory rearing of leopard darters (Percina pantherina). Final Report to U.S.
Army Corps of Engineers, Tulsa, Oklahoma. 27 pages.

Zale, A. V., V. H. Travnichek, and W. L. Fisher. 1990. Effects of the Pensacola

Hydropower Project on the fishery resource of the Grand River. Component A.1.
Seasonal downstream movements of ichthyoplankton past Pensacola Dam.
Final Report to Grand River Dam Authority, Vinita, Oklahoma. 56 pages.

Fisher, W. L., and A. V. Zale. 1990. Effects of the Pensacola Hydropower Project on
the fishery resource of the Grand River. Component A.2. Seasonal downstream
movements of juvenile and adult fish past Pensacola Dam. Final Report to
Grand River Dam Authority, Vinita, Oklahoma. 39 pages.

Zale, A. V., K. M. Sorenson, and W. L. Fisher. 1990. Effects of the Pensacola

Hydropower Project on the fishery resource of the Grand River. Component A.3.

26
 Grand Lake study: fish assemblage composition and entrainment susceptibilities.
Final Report to Grand River Dam Authority, Vinita, Oklahoma. 114 pages.

Zale, A. V., T. G. Adornato, and W. L. Fisher. 1990. Effects of the Pensacola

Hydropower Project on the fishery resource of the Grand River. Component C.
Effects on fisheries of establishment of a riverine environment in the headwaters
of Lake Hudson. Final Report to Grand River Dam Authority, Vinita, Oklahoma.
149 pages.

Fisher, W. L., and A. V. Zale. 1990. Effects of the Pensacola Hydropower Project on
the fishery resource of the Grand River. Component E. Effects of lake level
fluctuations resulting from power generation on littoral nursery habitats. Final
Report to Grand River Dam Authority, Vinita, Oklahoma. 31 pages.

Zale, A. V., H. VanOffelen, S. A. Bryan, E. Campbell, and E. Irwin. 1990. American

Fisheries Society student survey. Final Report to American Fisheries Society,
Bethesda, Maryland. 112 pages.

Zale, A. V., D. M. Leslie, Jr., W. L. Fisher, and S. G. Merrifield. 1989. The

physicochemistry, flora, and fauna of intermittent prairie streams: a review of the
literature. U.S. Fish and Wildlife Service Biological Report 89(5). 44 pages.

Zale, A. V., and S. G. Merrifield. 1989. Species profiles: life histories and
environmental requirements of coastal fishes and invertebrates (south Florida) --
tarpon and ladyfish. U.S. Fish and Wildlife Service Biological Report 82 (11.104).
U.S. Army Corps of Engineers, TR EL-82-4. 17 pages.

Zale, A. V., and S. G. Merrifield. 1989. Species profiles: life histories and
environmental requirements of coastal fishes and invertebrates (south Florida) --
reef-building tube worm. U.S. Fish and Wildlife Service Biological Report 82
(11.115). U.S. Army Corps of Engineers, TR EL-82-4. 12 pages.

Zale, A. V., D. M. Leslie, Jr., W. L. Fisher, and S. G. Merrifield. 1989. The role of
intermittent prairie streams in the dynamics of terrestrial and aquatic systems.
Final Report to U.S. Fish and Wildlife Service, Office of Information Transfer, Fort
Collins, Colorado. 129 pages.

Zale, A. V., and J. Burroughs. 1989. Water quality associated with the 1988 striped
bass mortality event at Keystone Reservoir, Oklahoma. Final Report.
Addendum to Federal Aid Project F-41-R (Oklahoma), Job XII. 8 pages.

Zale, A. V., and L. S. Jacks. 1988. Movements of striped bass in the Arkansas River
downstream from Keystone Reservoir. Final Report, Federal Aid Project F-41-R
(Oklahoma), Job XI. 22 pages.

27
Zale, A. V., R. L. Lochmiller, and J. D. Wiechman. 1988. Analysis of the suitability of
Keystone Reservoir, Oklahoma, for habitation by adult striped bass. Final
Report, Federal Aid Project F-41-R (Oklahoma), Job XII. 36 pages.

Bain, M., S. Meyers, and A. V. Zale. 1988. Surgical skin staplers may not be reliable
for biotelemetry research. U.S. Fish & Wildlife Service Research Information
Bulletin 88-67.

James, P. W., O. E. Maughan, and A. V. Zale. 1988. Reproductive ecology of the

leopard darter, Percina pantherina. Final Report, Endangered Species Project E-
2-R (Oklahoma), Job I. 27 pages.

Lechner, M., A. V. Zale, S. O'Donnell, and B. Ben. 1987. Distribution and abundances
of the leopard darter, Percina pantherina (Moore and Reeves) in the upper
Mountain Fork and upper Little River drainages. Final Report No. T860B19 to
Tulsa District, U.S. Army Corps of Engineers, Tulsa, Oklahoma. 27 pages.

Leon, S. C., M. T. Ferguson, O. E. Maughan, and A. V. Zale. 1987. Distribution and

abundances of the leopard darter, Percina pantherina (Moore and Reeves) in the
Saline, Cossatot, and Rolling Fork River drainages. Final Report to Tulsa
District, U.S. Army Corps of Engineers, Tulsa, Oklahoma. 27 pages.

Zale, A. V., and R. Mulholland. 1985. Habitat suitability index models: laughing gull.
U.S. Fish and Wildlife Service Biological Report 82(10.94). 23 pages.

Hingtgen, T. M., R. Mulholland, and A. V. Zale. 1985. Habitat suitability index models:
eastern brown pelican. U.S. Fish and Wildlife Service Biological Report
82(10.90). 20 pages.

Zale, A. V. 1985. Book review of W. R. Courtenay, Jr., and J. R. Stauffer, Jr., editors,
Distribution, Biology, and Management of Exotic Fishes. Transactions of the
American Fisheries Society 114:627-628. Invited

Montague, C. L., A. V. Zale, and H. F. Percival. 1985. A conceptual model of salt

marsh management on Merritt Island National Wildlife Refuge, Florida. Florida
Cooperative Fish and Wildlife Research Unit Technical Report No. 17. 92 pages.

Conrow, R., and A. V. Zale. 1985. Early life history stages of fishes of Orange Lake,
Florida: an illustrated identification manual. Florida Cooperative Fish and Wildlife
Research Unit Technical Report No. 15. 45 pages.

Montague, C. L., A. V. Zale, and H. F. Percival. 1984. Photographic analysis of natural

and impounded salt marsh in the vicinity of Merritt Island, Florida. Florida
Cooperative Fish and Wildlife Research Unit Technical Report No. 11. 23 pages.

28
Montague, C. L., A. V. Zale, H. F. Percival, and T. Hingtgen. 1984. A categorized
bibliography for a conceptual model of salt marsh management on Merritt Island,
Florida. Florida Cooperative Fish and Wildlife Research Unit Technical Report
No. 9. 98 pages.

Zale, A. V., C. L. Montague, H. F. Percival, and T. Hingtgen. 1984. Impounded coastal

wetlands of the Atlantic coast of the southeastern United States: a categorized
bibliography. Florida Cooperative Fish and Wildlife Research Unit Technical
Report No. 8. 93 pages.

Zale, A. V., and R. J. Neves. 1983. Freshwater mussels: a declining resource. Virginia
Wildlife 44(1):13-15.

Author or co-author of 236 technical presentations at scientific meetings, symposia,

workshops, and seminars since 1980; list available upon request.

29