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The Non-Vascular Plants

The non-vascular plants are divided into three taxonomic phyla: liverworts (Phylum
Hepatophyta), hornworts (Phylum Anthocerophyta), and mosses (Phylum Bryophyta). These groups
vary in a lot of ways, and may or may not be closely related. Evidence from the fossil record suggests that
they evolved during the Ordovician period. However, the incomplete fossil record makes it difficult to tell
how closely related the non-vascular plants are. Nevertheless, recent molecular data suggest that these
plants have individual lines of descent, and that they do not form a monophyletic group. The observation
that today’s mosses have a rudimentary form of vascular tissue suggests that they are more closely
related to vascular plants than the hornworts and liverworts.

Phylum Hepatophyta

General Characteristics:
Plants thallose or leafy, mostly bilaterally symmetrical, flattened, with isodiametric cells in the costate
leaves; sporophyte determinate, evanescent, and opening explosively into four valves.

Representative Specimens: Marchantia

Phylum Bryophyta

General Characteristics:
Plants leafy mostly radially symmetrical but, if flattened, with elongate cells or a costa in the leaves;
sporophyte determinate, persistent with a distinct cap which falls off preceding the release of spores.

Representative Specimen: moss

Note: Familiarize the parts and the function of each part.


The Spore-bearing Vascular Plants
Ferns, whisk ferns, horsetails, and lycophytes comprise a group of plants known as the spore-
bearing vascular plants. Like all land plants, they exhibit alternation of generations. These plants have a
large and prominent sporophyte with vascular tissue. Many have well-differentiated plant organs
(roots, stems, and leaves). The gametophyte generation in this group is small and lacks vascular tissue.
This group can be divided into two phyla: Lycophyta (lycophytes) and Pterophyta (whisk ferns,
horsetails, and ferns).

Phylum Pterophyta

Sori – it functions for reproduction

Horsetails
General Characteristics:
Small shrubs with conspicuously jointed, ribbed, unbranched stems, spore-bearing structures
grouped into strobili, leaves are classified as megaphyllous

Representative Specimen: Equisetum

Phylum Lycophyta

General Characteristics:
With true roots; leaves, called microphylls, are generally small, scale-like or needle-like, and
usually arranged spirally. Sporophylls (spore-bearing leaves) form strobili (singular, strobilus).

Representative Specimens: Lycopodium


Selaginella

Note: you should know how Lycopodium and Selaginella looks like.
The Gymnosperms
The gymnosperms, represented by pines, firs, spruces, and cycads, among others, mostly have
well-developed cones bearing unprotected seeds on the surface of the scales. Their seeds are
not enclosed in fruits, hence their name (Greek, gymnos, naked, and sperma, seed). There are
about 780 gymnosperm species, which are more of temperate distribution. Only two
taxonomic phyla (Cycadophyta and Coniferophyta) will be covered in the laboratory due to the
availability of specimens.

Phylum Cycadophyta

General Characteristics:
Dioecious plants with palm-like or fernlike forms; strobili (singular, strobilus) present; motile
sperms present.

Representative Specimen: Cycas

Note: In addition, you should also know the structure of the Cycas
Female cone

The Angiosperms
Flowering plants, or angiosperms, belong to the Phylum Anthophyta. Angiosperms are a very
large and successful group, consisting of at least 300,000 known species. Flowering plants are
named angiosperms (Greek, angion, vessel, and sperma, seed), because unlike the
gymnosperms, their ovules are always enclosed within diploid tissues. The term angio in their
name refers to the ovary, which develops into the fruit – a unique characteristic of this group.

Most members of Phylum Anthophyta belong to one of two large taxonomic classes. These
classes are the Monocotyledones (or Liliopsida) and the Dicotyledones (or Magnoliopsida).

Class Monocotyledones / Liliopsida

Characteristics: With one cotyledon, leaves usually parallel-veined, floral parts (except carpels)
in sets of 3’s or 4’s, root system usually adventitious.

1. Family Orchidaceae

Vegetative characters: epiphytic, velamen present (note: white roots); leaves parallel-veined
Reproductive characters: inflorescence solitary or racemes; flowers irregular: innermost
enlarged (labellum/lip); stamens united to form a column (gynandrium)
Note: Familiarize the classification From Kingdom until Family.

Class Dicotyledones / Magnoliopsida

Characteristics: Two cotyledons; leaves usually net-veined; 50% are woody.

1. Family Cucurbitaceae

Vegetative characters: tendril-bearing vines with 1 tendril/node); leaves spirally alternate


Reproductive characters: flowers solitary or in axillary inflorescences; flowers unisexual, regular
(usually); ovary inferior; placentation usually parietal; fruit a pepo (usually)
Specimens: Cucurbita pepo (squash, pumpkin), Momordica charantia (ampalaya/bitter gourd)

2. Family Asteraceae / Compositae

Vegetative characters: habit herbs (mostly), shrubs or vines; leaves alternate, simple or compound
Reproductive characters: inflorescences heads; flowers bisexual or unisexual, regular to irregular; ovary
inferior; fruit an achene (usually)
Specimens: Helianthus (sunflower), Chrysanthemum, Cosmos

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