Lopingian

Lopingian
Lopingian
259.51 ± 0.21 – 251.902 ± 0.024 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	A map of Earth as it appeared 255 million years ago during the Lopingian Epoch, Wuchiapingian Age
Chronology
←	Permian-Triassic mass extinction event
←	end-Capitanian extinction event
←	Olson's Extinction
	Subdivision of the Permian according to the ICS, as of 2023.; Vertical axis scale: Millions of years ago
Etymology
Name formality	Formal
Synonym(s)	Late/Upper Permian;
Usage information
Celestial body	Earth
Regional usage	Global (ICS)
Time scale(s) used	ICS Time Scale
Definition
Chronological unit	Epoch
Stratigraphic unit	Series
Time span formality	Formal
Lower boundary definition	FAD of the Conodont Clarkina postbitteri postbitteri
Lower boundary GSSP	Penglaitan Section, Laibin, Guangxi, China; 23.6953°N 109.3211°E
Lower GSSP ratified	2004
Upper boundary definition	FAD of the Conodont Hindeodus parvus.
Upper boundary GSSP	Meishan, Zhejiang, China; 31.0798°N 119.7058°E
Upper GSSP ratified	2001

The Lopingian is the uppermost series/last epoch of the Permian.^[4] It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic.

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The Lopingian is often synonymous with the informal terms late Permian or upper Permian.

The name was introduced by Amadeus William Grabau in 1931 and derives from Leping, Jiangxi in China.^[5] It consists of two stages/ages. The earlier is the Wuchiapingian and the later is the Changhsingian.^[6]

The International Chronostratigraphic Chart (v2018/07)^[4] provides a numerical age of 259.1 ±0.5 Ma. If a Global Boundary Stratotype Section and Point (GSSP) has been approved, the lower boundary of the earliest stage determines numerical age of an epoch. The GSSP for the Wuchiapingian has a numerical age of 259.8 ± 0.4 Ma.^[7]^[8]

Evidence from Milankovitch cycles suggests that the length of an Earth day during this epoch was approximately 22 hours.^[9]

During the Lopingian, most of the earth was in the supercontinent Pangaea. The Zechstein sea, would, at times, be connected to the Paleotethys; Other features of the earth during the time were the Microcontinent Cathaysia; And the Cimmerian superterrane, which divided the Tethys Ocean realm into the Paleo-Tethys Ocean and the slowly expanding Neotethys Ocean.

The Lopingian ended with the Permian–Triassic extinction event, where over 95% of species went extinct.

The series follows the Guadalupian, which ended with the Capitanian mass extinction, during which many species of brachiopods, ammonoids and other groups went extinct.^[10]

Conodonts would reach their all-time low during this period, despite this, they are recovered from most marine Permian localities.^[11] Common conodonts from the Lopingian include the genera Clarkina and Hindeodus.

The Lopingian would see the decline of the Paleozoic ammonoid orders (Goniatitida and Prolecanitida) and the rise of the order Ceratitida, especially within the superfamily Xenodiscoidea.^[12]

Only seven trilobites are known from the Lopingian, with only five by the end of the epoch. One of the last members of this clade was Kathwaia capitorosa.^[13]

Eurypterids were nearly extinct by this point, consisting of the possibly Lopingian Campylocephalus permicus of Russia; and the Changhsingian Woodwardopterus? freemanorum of Australia.^[14]

A member of the extant Horseshoe crab family, Limulidae; Guangyuanolimulus appears at the end of the period.^[15]

On land, gorgonopsians would become the apex predators after the extinction of the Dinocephalians, Other predators include the Therocephalians. Herbivorous animals of the Lopingian include the pareiasaurs such as Scutosaurus or dicynodonts, such as Dicynodon.

Fossil gallery

Smilesaurus ferox, one of the most fearsome predators of the Lopingian
Skull of Dicynodon, a dicynodont.
Scutosaurus, a common herbivore during the late Permian
Paratirolites, an ammonoid from the latest Permian (Changhsingian)

Geologic time scale – System that relates geologic strata to time
Cisuralian – First series of the Permian

[1]
"International Chronostratigraphic Chart" (PDF). International Commission on Stratigraphy. September 2023. Retrieved 16 December 2024.
[2]
Jin, Yugan; Shen, Shuzhong; Henderson, Charles; Wang, Xiangdong; Wang, Wei; Wang, Yue; Cao, Changqun; Shang, Qinghua (December 2006). "The Global Stratotype Section and Point (GSSP) for the boundary between the Capitanian and Wuchiapingian Stage (Permian)" (PDF). Episodes. 29 (4): 253–262. doi:10.18814/epiiugs/2006/v29i4/003. Retrieved 13 December 2020.
[3]
Hongfu, Yin; Kexin, Zhang; Jinnan, Tong; Zunyi, Yang; Shunbao, Wu (June 2001). "The Global Stratotype Section and Point (GSSP) of the Permian-Triassic Boundary" (PDF). Episodes. 24 (2): 102–114. doi:10.18814/epiiugs/2001/v24i2/004. Retrieved 8 December 2020.
[4]
International Commission on Stratigraphy. "Chart". Retrieved 10 July 2018.
[5]
Zhang, Shouxin (2009). Geological Formation Names of China (1866–2000). Beijing/Dordrecht: Higher Education Press/Springer. p. 681. ISBN 978-7-040-25475-4.
[6]
Allaby, Michael (2015). A Dictionary of Geology and Earth Sciences (4th ed.). Oxford University Press. doi:10.1093/acref/9780199653065.001.0001. ISBN 9780199653065.
[7]
International Commission on Stratigraphy. "GSSPs". Retrieved 10 July 2018.
[8]
Gradstein, Felix M.; Ogg, James G.; Smith, Alan G. (2004). A Geologic Time Scale 2004. Cambridge University Press. ISBN 9780521786737.
[9]
Wu, Huaichun; Zhang, Shihong; Hinnov, Linda A.; Jiang, Ganqing; Feng, Qinglai; Li, Haiyan; Yang, Tianshui (13 September 2013). "Time-calibrated Milankovitch cycles for the late Permian". Nature Communications. 4: 2452. Bibcode:2013NatCo...4.2452W. doi:10.1038/ncomms3452. PMC 3778519. PMID 24030138. S2CID 6477972.
[10]
Bond, David; Hilton, Jason; Wignall, Paul; Ali, Jason; Stevens, Liadan; Sun, Yadong; Lai, Xulong (2010). "The Middle Permian (Capitanian) mass extinction on land and in the oceans". Earth-Science Reviews. 102 (1–2): 100–116. Bibcode:2010ESRv..102..100B. doi:10.1016/j.earscirev.2010.07.004.
[11]
Wardlaw, R. B. (1995). "Permian Conodonts". In Scholle, P.A.; Peryt, T.M.; Ulmer-Scholle, D.S. (eds.). The Permian of Northern Pangea. Berlin, Heidelberg: Springer. pp. 186–195. doi:10.1007/978-3-642-78593-1_12. ISBN 978-3-642-78595-5.
[12]
Leonova, T.B. (2016). "Major trends in the evolution of Permian ammonoids". Paleontological Journal. 50 (2): 131–140. Bibcode:2016PalJ...50..131L. doi:10.1134/S0031030116020039.
[13]
"The last Trilobites". www.Trilobites.info.
[14]
Poschmann, Marjus J.; Rozefelds, Andrew (2022). "The last eurypterid – a southern high-latitude record of sweep-feeding sea scorpion from Australia constrains the timing of their extinction". Historical Biology. 34 (10): 2020–2030. Bibcode:2022HBio...34.2020P. doi:10.1080/08912963.2021.1998033.
[15]
Hu, Shixue; Feldmann, Rodney M.; Schweitzer, Carrie E.; Benton, Michael J.; Huang, Jinyuan; Wen, Wen; Min, Xiao; Zhang, Qiyue; Zhou, Changyong; Ma, Zhixin (15 September 2022). "A new horseshoe crab from the Permian-Triassic transition of South China: Xiphosurids as key components of post-extinction trophic webs". Palaeogeography, Palaeoclimatology, Palaeoecology. 602: 111178. doi:10.1016/j.palaeo.2022.111178. ISSN 0031-0182.

[ICS-1] [1]
"International Chronostratigraphic Chart" (PDF). International Commission on Stratigraphy. September 2023. Retrieved 16 December 2024.

[2] [2]
Jin, Yugan; Shen, Shuzhong; Henderson, Charles; Wang, Xiangdong; Wang, Wei; Wang, Yue; Cao, Changqun; Shang, Qinghua (December 2006). "The Global Stratotype Section and Point (GSSP) for the boundary between the Capitanian and Wuchiapingian Stage (Permian)" (PDF). Episodes. 29 (4): 253–262. doi:10.18814/epiiugs/2006/v29i4/003. Retrieved 13 December 2020.

[3] [3]
Hongfu, Yin; Kexin, Zhang; Jinnan, Tong; Zunyi, Yang; Shunbao, Wu (June 2001). "The Global Stratotype Section and Point (GSSP) of the Permian-Triassic Boundary" (PDF). Episodes. 24 (2): 102–114. doi:10.18814/epiiugs/2001/v24i2/004. Retrieved 8 December 2020.

[Chart-4] [4]
International Commission on Stratigraphy. "Chart". Retrieved 10 July 2018.

[5] [5]
Zhang, Shouxin (2009). Geological Formation Names of China (1866–2000). Beijing/Dordrecht: Higher Education Press/Springer. p. 681. ISBN 978-7-040-25475-4.

[6] [6]
Allaby, Michael (2015). A Dictionary of Geology and Earth Sciences (4th ed.). Oxford University Press. doi:10.1093/acref/9780199653065.001.0001. ISBN 9780199653065.

[7] [7]
International Commission on Stratigraphy. "GSSPs". Retrieved 10 July 2018.

[a2004-8] [8]
Gradstein, Felix M.; Ogg, James G.; Smith, Alan G. (2004). A Geologic Time Scale 2004. Cambridge University Press. ISBN 9780521786737.

[WuEtAl2013-9] [9]
Wu, Huaichun; Zhang, Shihong; Hinnov, Linda A.; Jiang, Ganqing; Feng, Qinglai; Li, Haiyan; Yang, Tianshui (13 September 2013). "Time-calibrated Milankovitch cycles for the late Permian". Nature Communications. 4: 2452. Bibcode:2013NatCo...4.2452W. doi:10.1038/ncomms3452. PMC 3778519. PMID 24030138. S2CID 6477972.

[10] [10]
Bond, David; Hilton, Jason; Wignall, Paul; Ali, Jason; Stevens, Liadan; Sun, Yadong; Lai, Xulong (2010). "The Middle Permian (Capitanian) mass extinction on land and in the oceans". Earth-Science Reviews. 102 (1–2): 100–116. Bibcode:2010ESRv..102..100B. doi:10.1016/j.earscirev.2010.07.004.

[11] [11]
Wardlaw, R. B. (1995). "Permian Conodonts". In Scholle, P.A.; Peryt, T.M.; Ulmer-Scholle, D.S. (eds.). The Permian of Northern Pangea. Berlin, Heidelberg: Springer. pp. 186–195. doi:10.1007/978-3-642-78593-1_12. ISBN 978-3-642-78595-5.

[12] [12]
Leonova, T.B. (2016). "Major trends in the evolution of Permian ammonoids". Paleontological Journal. 50 (2): 131–140. Bibcode:2016PalJ...50..131L. doi:10.1134/S0031030116020039.

[13] [13]
"The last Trilobites". www.Trilobites.info.

[14] [14]
Poschmann, Marjus J.; Rozefelds, Andrew (2022). "The last eurypterid – a southern high-latitude record of sweep-feeding sea scorpion from Australia constrains the timing of their extinction". Historical Biology. 34 (10): 2020–2030. Bibcode:2022HBio...34.2020P. doi:10.1080/08912963.2021.1998033.

[15] [15]
Hu, Shixue; Feldmann, Rodney M.; Schweitzer, Carrie E.; Benton, Michael J.; Huang, Jinyuan; Wen, Wen; Min, Xiao; Zhang, Qiyue; Zhou, Changyong; Ma, Zhixin (15 September 2022). "A new horseshoe crab from the Permian-Triassic transition of South China: Xiphosurids as key components of post-extinction trophic webs". Palaeogeography, Palaeoclimatology, Palaeoecology. 602: 111178. doi:10.1016/j.palaeo.2022.111178. ISSN 0031-0182.

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Lopingian

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References