Catalogue of The World Gonocephalum
Catalogue of The World Gonocephalum
Catalogue of The World Gonocephalum
1
Museum and Institute of Zoology Polish Academy of Sciences, Wilcza 64,
00-679 Warszawa, Poland; e-mail: [email protected]
2Department of Entomology, The Swedish Museum of Natural History,
Abstract.— The catalogue of all known World species of the genus Gonocephalum Solier,
1834 is presented: 3 subgenera, 415 species and subspecies are listed. The data of primary
types and distribution for known species and subspecies are included. Gonocephalum
zoltani nom. nov. is proposed as replacement name for Gonocephalum parallelum
Kaszab, 1952 (nec Gonocephalum setulosum var. parallelum Normand, 1936) which is
junior homonym; Gonocephalum neoblairi nom. nov. is proposed as replacement name
for Gonocephalum Blairi Kaszab, 1952 (nec Opatrum blairi Gebien, 1922, presently
placed in Gonocephalum) which is junior homonym. The following new homonyms are
presented for the species presently placed in Gonocephalum: Hopatrum arenarium
Fairmaire, 1894 nom. preoccup., nec Opatrum arenarium Fabricius, 1775; Gonoce-
phalum curvicolle Gridelli, 1934 nom. preoccup., nec Gonocephalum curvicolle Reitter,
1889; Gonocephalum pubens Gebien, 1913 nom. preoccup., nec Opatrum pubens
Marseul, 1876; Opatrum pusillum Küster, 1849 nom. preoccup., nec Opatrum pusillum
Fabricius, 1791; Opatrum viennense Küster, 1849 nom. preoccup., nec Gonocephalum
viennense Duftschmid, 1812; Opatrum modestum Küster, 1849 name preoccup. nec
Opatrum modestum Reiche, 1847; Opatrum patruele Küster, 1849 nom. preoccup. nec
Opatrum patruele Erichson, 1843; Opatrum setulosum Küster, 1849 nom. preoccup.,
nec Opatrum setulosum Faldermann, 1837. Incertae sedis taxa are listed separately.
Gonocephalum acoriaceum Chûjô, 1975: 15. – Iwan and Löbl 2008: Opatrum adpressum Germar, 1824: 145.
262. Hopatrum adpressum Germar, 1824. – Gemminger, 1870: 1930.
Gonocephalum adpressum (Germar, 1824). – Gebien, 1910b: 322,
1913: 376; 1939: 448; Kaszab 1952a: 652.
Type data. Holotype: male (FAKU No. 2044), “Mt.
Opatrum complanatum Guérin-Méneville, 1830: 98.
Banna, Ishigaki I., the Ryukyus, at light, 24.V.1975, Hopatrum complanatum Guérin-Méneville, 1830. – Gemminger,
H. Makihara leg.”. 1870: 1931.
Paratypes: Kurio, Is. Yaku-shima, 3.VII.1959, M. et Gonocephalum complanatum (Guérin-Méneville, 1830). – Boisduval,
T. Okadome leg., 1 male (FAKU); Kukibachi, Tane- 1835: 252; Gebien, 1910b: 322, 1939: 448; Kaszab, 1952a: 683.
Gonocephalum alticola seychellense Ferrer, 2000b: 106. – Ferrer G. amplithorax (Fairmaire, 1894)
2004: 90.
Opatrum amplithorax Fairmaire, 1894: 659. – Champion 1895: 115.
Gonocephalum amplithorax (Fairmaire, 1894: 659). – Gebien 1910b:
Type data. Holotype: male (MRAC), “Seychelles,
322; Gridelli 1940 (1939): 240 (syn. Gonocephalum simplex
Mahé Sud, Anche à la Mouche, 16–25.VII.1972, P.L.G. (Fabricius, 1801); Ferrer 1993a: 95 (bona species); Ferrer 2000b:
Benoit, J.J. Van Moi leg.”. 71, 2004: 88.
Paratypes: Seychelles, Mahé Sud, Anche à la
Mouche, 16–25.VII.1972, P.L.G. Benoit, J.J. Van Moi Type data. Holotype, male (MNHN), “Abyss. Raf-
leg., 5 males, 7 females (MRAC), 2 ex. (JFCS); Sey- fray”.
chelles 2 ex. (BMNH); Mahé, Mamella, A. Brauer leg., Distribution. AFROTROPICAL REGION: Erythrea.
32 ex. (ZMHB); Andasibé, Perinet, 930–1.000 m
6–8.II.1993, G. Dunay, J. Janàk leg., 70 ex. (ZSMB);
Ranomafana, 11.V.1989, L. Bartolozzi, S. Taiti, C. G. andamanense Kaszab, 1952
Raharimina, E. Randrianosolo leg., 3 ex. (MZUF 7809),
Gonocephalum andamanense Kaszab, 1952a: 539.
1 ex. (JFCS); Ambanja, Benanemiky-Morovato, 21.IX.
1989, L. Bartolozzi, S. Taiti, C. Raharimina leg., 2 ex.
(MZUF 7810); Tsimbazaza, Antananarivo, 7.XII.1989, Type data. Holotype (NHMB).
id., 1 ex. (MZUF 7811); Sambirano, Andimanka, 20.IX. Material studied. “Andaman-Insel (leg. Wimberley)”.
Other depository. BMNH (2 ex.)
1989, 1 ex. (MZUF 7812); Perinet, 7–8.IX.1989, id., 1 ex.
Distribution. ORIENTAL REGION: Andaman Isl.
(MZUF 7813); Massif d’Ambondrombe (Madagascar
Est) 1.200–1.300 m., env. d’Ikoka, crête d’Ambroasa,
camo 1, 9–10.III.1996, Janak, P. Moravec leg. (forêt
G. andrewesi Kaszab, 1952
humide, sous écorces) 1 ex. (ZSMG); Mayotte, 1884,
L. Humboldt leg., 35 ex. (MNHN), 1 ex. (JFCS); Sey- Gonocephalum Andrewesi Kaszab, 1952a: 649. – Chûjô 1968: 21.
chelles, ex coll. L. Fairmaire, 1 ex. (MNHN). Gonocephalum andrewesi Kaszab 1952a. – Hau Li 2002: 138; Iwan
Distribution. AFROTROPICAL REGION: Mada- and Löbl 2008: 262.
gascar (Ankaratra, Erakoka), Seychelles.
Type data. Syntypes: 5 ex. (SDEI).
Material studied. Christmas-Insel, I–IV.1933; S. O.
G. alticola zulu Ferrer, 2000 Borneo (leg. Grabowsky); Sarawak, Brooketon (leg. D.
S. Mjöberg); Brit. N. Borneo, Labuan Id. (leg. Dr. E.
Gonocephalum alticola zulu Ferrer, 2000b: 105. Mjöberg); Buitenzoorg (leg. M. Fleischer); Nongkodjad-
jar, 4000 ft (leg. J. P. A. Kalis); Sindanglaya, 1870 (leg.
Type data. Holotype: male (TMNH), “Afrique du J. Xántus); Preanger Reg., Tjisorve pan., 4300 ft, 24.IV.
Sud: Transvaal, Zululand, Sordwana Bay, 10.XI.1984, 1923 (leg. H. M. Pendlebury); Tjibodas, 1400 m, VIII.
Scholtz et Bellamy leg.”. 1921; Semongkat am Nordhang der Batoe Lanteh
Paratypes: Afrique du Sud: Transvaal, Zululand, Gebirge (leg. Rensch); Sumbawa, Dempoe (leg. Ren-
Sordwana Bay, 10.XI.1984, Scholtz et Bellamy leg., sch); Timor, Atapupu (leg. v. Martens); Lombok, Sapit,
femelle (TMNH), 1 ex. (JFCS); Mtunzini, 30.XI.1963, 2000 ft (leg. H. Fruhstorfer); Lombok, Sumbalun, 4000
L. Vari leg., 2 ex. (TMNH); Durban, 3.I.1966, M. Webb ft (leg. H. Fruhstorfer); Bali, Selat; Flores (coll.
leg., 2 ex. (TMNH); Zululand, XI.1905, I. Trägårdh Fry); Ceram Küste (leg. E. Streesemann); Key-Insel,
leg., 3 ex.; Pinetown, II.1896, Cragoe leg., 2 ex. Gu. Daab, 300 m (leg. H. C. Siebers); Tual (leg. C.
(MNHN). Ribbe); Roma-Insel; Larat-Insel (=Tenimber Insel);
250 D. IWAN, J. FERRER and M. RAŚ
Type data. Lectotype (designated by Ferrer 2000b: Type data. Holotype: female (ZMHB), “Mozam-
113): male (ZMHB), “Angola, Schonlein leg. (47938) typ!”. bique, Tete, Peters leg.”, designe comme lectotype;
Paralectotype: Angola, Schonlein leg. (47938) typ!, Distribution. AFROTROPICAL REGION: Mada-
male (ZMHB). gascar, Mozambique, Republic of South Afrika, Soma-
Distribution. AFROTROPICAL REGION: Angola lia, Zimbabwe.
(Namib Desert). Remarks. Gonocephalum angusticolle (Ger-
staecker) determined by Español and Viñolas (1983:
36) belongs to Gonocephalum indictum Ferrer, 1995.
G. angolense escalerai Ferrer, 2000
Gonocephalum apterum Koch, 1955a: 528. – Ferrer 1993a: 74, G. asperatum Gebien, 1920
1995a: 6.
Gonocephalum asperatum Gebien, 1920: 232. – Gebien 1939: 450;
Kaszab 1952a: 557.
Type data. Holotype: (TMNH), “South-central Cape
Province: Zwartkops, Algoa Bay, November 1921, H.
Brauns”. Type data. Unknown.
Distribution. AUSTRALIAN AND PACIFIC RE-
Allotype: (TMNH), “South-central Cape Province:
GIONS: Australia, New Guinea.
Zwartkops, Algoa Bay, November 1921, H. Brauns”.
Paratypes: South-central Cape Province: Zwart-
kops, Algoa Bay, November 1921, H. Brauns., 4 ex.
G. assimile (Küster, 1849)
(TMNH); Port Elizabeth, C. G. C. Dicson, 2 ex. (TMNH).
Distribution. AFROTROPICAL REGION: Republic of Opatrum assimile Küster, 1849a: 52.
South Africa (Cape Province). Hopatrum assimile Küster, 1849. – Gemminger 1870: 1930.
252 D. IWAN, J. FERRER and M. RAŚ
Gonocephalum assimile (Küster, 1849). – Baudi 1876 (1875?): 67, G. bartolozzii Ferrer, 1995
(232?); Seidlitz 1894: 439; Reitter 1904: 144; Gebien 1910b: 322,
1939: 444; Portevin 1934: 13; Schuster 1928: 989; Reichardt 1936: Gonocephalum bartolozzii Ferrer, 1995a: 23. – Ferrer 1995b: 40,
102; Espanol 1952: 5; Gardini 1975: 685; Canzoneri 1977: 274; Can- 1998a: 40;
zoneri and Vienna 1987: 20; Ferrer 1993a: 76, 1995a: 5; Iwan and
Löbl 2008: 262.
Type data. Holotype: male (MZUF)(HT 7824),
“Somalie, Bud Bud, 15.VIII.1968 (S.B.S.)”.
Type data. „In Sardinien. Beschreiben nach dem
Paratypes: Somalie: Karora, I–III.1938, «Gono-
Originalexemplar der Sturm’schen Sammlung“. cephalum – Gridelli det. 1941», PT 7825: 1 female
Distribution. PALAEARCTIC REGION (Iwan and (MZUF); PT 7841: Ola Uager, Chisimaio, 14.IX.1964,
Löbl 2008): Europe: Italy (incl. Sardinia, Sicily, San Miss. Biol. Somalia 1964, 1 female (MZUF); PT 7826:
Marino). Afmadu, 8.IX.1964, Miss. Biol. 64, PT 7826: 1 female
Remarks. France (Canzoneri and Vienna 1987: (MZUF); PT 7842: km 39.48 O Garoe/Gardo, 4.X.1977
20). (S.B.S.) 3 ex. (MZUF), 2 ex. (JFCS).
Distribution. AFROTROPICAL REGION: Somalia.
Gonocephalum Balmeae Kaszab, 1952a: 541. Gonocephalum bengalense Kaszab, 1952a: 637.
Type data. Syntypes: 2 ex. (BMNH), “Satara Distr., Type data. Syntypes: 5 ex. (NHMB), “Bengalen;
3700 ft, 23.IV.1912 (leg. F. H. B.)”. Swada”.
Distribution. ORIENTAL REGION: India. Distribution. ORIENTAL REGION: India.
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 253
G. besnardi besnardi Kaszab, 1982 Gonocephalum bilineatum (Walker, 1858). – Gebien 1910b: 322,
1921: 269, 1939: 446; Chatanay 1917: 238; Blair 1921: 269; Schuster
Gonocephalum besnardi Kaszab, 1982a: 163. – Ferrer 1995a: 58 1928: 989; Reichardt 1936: 109; Kaszab 1952a: 641, 1954: 75, 1965a:
(stat. subsp.); Kaszab 1982b: 163; Schawaller 2010: 268; Iwan and 114, 1965d: 287, 1970a: 424, 1977: 257, 1979a: 260, 1979b: 65, 1980b:
Löbl 2008: 262. 171; Boddy 1965: 171; Kwon and Choi 1986: 106; Schawaller 1997a:
3; Hua Li 2002: 138; Iwan and Löbl 2008: 263.
Gonocephalum Kamtschaticum Motschulsky, 1860: 139.
Type data. Holotype: male (ZSMG), “Ver. Arab.
Hopatrum kamtschaticum Motschulsky, 1860. – Gemminger 1870:
Emirate, Ras Al-Khayma, 24.II.1978, Cl. Besnard”. 1932.
Paratypes: Ver. Arab. Emirate, Ras Al-Khayma, Gonocephalum kamtschaticum Motschulsky, 1860. – Reitter 1904:
24.II.1978, Cl. Besnard, 15 ex. (ZSMG, TTM); Arabia 146; Gebien 1910b: 323; Schuster 1928: 989; Reichardt 1936: 109;
[Oman], Aiyum, 5.III.1947, W. Thesiger, 2 ex. (BMNH). Kaszab 1952a: 682, 1965d: 287.
Blapstinus latifrons LeConte, 1874: 70. – Casey 1890: 393;
(Ferrer 1995a: 58) Couple comparée a l’holotype (Ras
Gonocephalum latifrons (LeConte, 1874). – Champion 1895: 115;
el Khaima) par le Dr. O. Mérkl (HNHM). Gebien 1910b: 323, 1939: 452; Kaszab 1952a: 682; Ferrer 2000: 129.
Distribution. PALAEARCTIC REGION (Iwan and Opatrum orarium Lewis, 1894: 380. – Champion 1895: 117.
Löbl 2008): Asia: Arab Emirates; Oman; Saudi Arabia. Gonocephalum orarium (Lewis, 1894). – Gebien 1910b: 324;
Remarks. Afghanistan (Kaszab 1982b). Reichardt 1936: 109; Schuster 1928: 989; Kaszab 1952a: 682.
Type data. Holotype: male (MNHN), “Iran: Shadat, Gonocephalum bimaculatum Ferrer, 1995a: 61.
Lut, Iran, 12.III.1965, F. Pierre leg.”.
Paratypes: Iran, Iranshaar, 14.IV.1965, Mission Type data. Holotype: male (HNHM), “S. Afrique:
franco-iranienne, 3 females (MNHN at Ferrer coll.). Natal, 1924, «bimaculatum n. sp. Blair det. (in litt.).”.
Distribution. PALAEARCTIC REGION (Iwan and Paratypes: S. Afrique: Natal, 1924, «bimaculatum n.
Löbl 2008): Asia: Iran. sp. Blair det. (in litt.), 1 male, 2 females (HNHM); Trans-
vaal: «Caffraria», J. Wahlberg leg. (1838?) «O. magicum
Bhm» (Boheman det. in litt.), 3 ex. (TMNH); Cape Prov.:
G. bigranulatum Kaszab, 1952 28 km SW of Barkly East, II.1967, L. Schultze leg., 6 ex.
(TMNH); Rodhes, 10.III.1951, 3 ex.Brinck and Rude-
Gonocephalum bigranulatum Kaszab, 1952a: 491.
beck, 1 ex. (TMNH); NE of Barkly East, 8–9.III.1992,
H.J. Bremer leg. 20 ex. (ZSMB), 3 ex. (JFCS); Her-
Type data. Holotype: single specimen (HNHM), schell, 1 ex. (TMNH); Springfontein, XII.1947–I.1948,
“Madras, Ramandroog”. P. Jackson leg., 2 ex. (TMNH); Koosfontein, XII.1908,
Distribution. ORIENTAL REGION: India. S. Viljoen leg., 1 ex. (TMNH); Reitz, 16.VII.1977, Mar-
querd leg., 1 ex. (HNHM); Lesotho: Nasareth, 20 miles
ESE Maseru, 26.III.1951, Brinck and Rudebeck leg.,
G. bilineatum (Walker, 1858)
1 ex. (ZMUL); Mt Machache, 25 miles E Maseru, 1951,
Opatrum bilineatum Walker, 1858: 284. id., 2 ex. (ZMUL); Makheke Mts, 10 miles ENE Mokoth-
Hopatrum bilineatum Walker, 1858. – Gemminger 1870: 1931. long, 7.IV.1951, id., 29 ex. (ZMUL); 4 ex. (JFCS);
254 D. IWAN, J. FERRER and M. RAŚ
Type data. Syntypes: 5 ex. (HNHM). Gonocephalum brachelytra Kaszab, 1952a: 632. – Kaszab 1968b: 20,
Material studied. “Astrilabe Bay, Stephansort, 1897 1970c: 152; Iwan and Löbl 2008: 263.
Paratypes: Kenya or. Malindi, 13–29.VIII.1983, H.J. Transvaal prov. Ermelo, 26.III.1985, no. 2048, T.W.
Bremer leg., 4 ex. (HNHM), 9 ex. (ZSMB), 1 ex. (JFCS); Drinkwater leg., 3 ex.; id. 11.IV. no. 2567 (1) id. 18.IV.
Lake Baringo Lodge, 31.III.1988, R. Grimm leg., 1 ex. no. 2037, 5 ex., et no. 2048, 4 ex.; id. 30.III.1986, no.
coll. R. Grimm; Zanzibar, Hildbrandt, 2664, 1 ex. 2596, 1 ex.; id. 12.III, no.2513, 13 ex., no.2515, 1 ex. et
(ZMHB); Tanzanie: Île de Zanzibar, Bububu, Alluaud no.591, 1 ex. (ZMUB); Johannesburg, III.1953, G. Rude-
and Jeannel, IV.1912, 21 ex. (MNHN); 3 ex. (JFCS). beck, III.1953, 4 ex. (TMNH); Lydenburg, P.A. Kraatz
Distribution. AFROTROPICAL REGION: Kenya, leg. 1896, 1 ex. (TMNH); Orange Free State: Bothaville,
Tanzania (Zanzibar). 18.IV.1979, T.W. Drinkwater leg. no. 1209, 2 ex.; id.
4.IV, no. 1221, 2 ex.; id. 25.IV; no. 1260, 1 ex.; id. 6.IV.
no.1323, 1 ex.; Clocolan, 23.IV.1985, no. 2070, 5 ex.;
G. brenieri Chatanay, 1917 Dewotsdorp; 26.III, no.2008; Exxelsior, 3.IV, no. 2215,
1 ex.; Frankfort, 3.I.1974, no. 1119, 1 ex.; Klerksdorp,
Gonocephalum Brenieri Chatanay, 1917: 238. – Gebien 1939: 448;
Kaszab 1952a: 659. 27.III.1977, n°1049, 4 ex.; 24.III.1983, no. 1566, 5 ex., et
no. 1569, 2 ex.; Ladybrand, 18.IV.1985, no. 2037, 5 ex.;
Type data. Syntypes: “Song Kong, coll. Chatanay” id. no. 2077 2 ex.; id. 23.IV, no.2181, 3 ex.; Lichtenburg,
(2 ex. MNHN). 12.V.1985, no. 1746, 1 ex.; Marquard, 23.IV.1985, no.
Distribution. ORIENTAL REGION: Vietnam. 2048, 4 ex., no. 2089; id. 16.IV, no. 2062, 1 ex.; Thenuis-
sen, 25.III. 1985, no. 1894, 1 ex.; id. 16.IV, no. 2062, 1
ex.; Virginia, 3.IV.1985, no. 2208, 1 ex.; id. 25.IV, no.
G. brevicorne Chatanay, 1917 2143, 1 ex.; Wesselsbron, 17.I.1979, no. 1138, 1 ex.; id.
11.IV.1979, no. 1241, 1 ex., et no. 1248, 2 ex. (ZMUB); 14
Gonocephalum brevicorne Chatanay, 1917: 247. – Blair 1928: 23;
km W Harrismith, XI.1963, M.L. Penrih lrg., 1 ex.; Farm
Gebien 1939: 448; Kaszab 1952a: 500; Kaszab and Chûjô 1966: 53.
Die River (11 miles S Ficksburg, XII. 1970, R. Joens,
K.M. Mathabathe leg., 14 ex. (ZMUB).
Type data. Syntypes: “Martapura, S.E de Borneo,
Distribution. AFROTROPICAL REGION: Lesotho,
1891, Doherty, coll. Chatanay” (MNHN).
Republic of South Africa (Transvaal, Orange Free State).
Distribution. AUSTRALIAN AND PACIFIC RE-
GIONS: Australia. ORIENTAL REGION: India, Indo-
nesia (Borneo, Java, Krakatau, Molukken, Sumatra), G. brittoni Kaszab, 1952
Malaysia, Thailand.
Gonocephalum Brittoni Kaszab, 1952a: 618.
Gonocephalum brincki Ferrer, 2000b: 107. – Ferrer 2004: 90. Type data. Syntypes: 1 ex. (NHMB), 1 ex. (HNHM),
“Java, Buitenzorg”, “Sumatra, leg. Dr. Machik”.
Type data. Holotype: male (ZMUL), “Lesotho: Distribution. ORIENTAL REGION: Indonesia
Nazareth MS; 20 miles SE Maseru, 24.III.1951, P. (Java, Sumatra).
Brinck, G. Rudebeck leg.”
Paratypes: Mt Machache, 25 miles E Maseru,
20.III.1951, id., 1 ex. (JFCS); Mamathes, 5 miles NE G. caesum Steven, 1829
Teytenayeng, 20.III.1951, id., 1 ex. (ZMUL); id. III.1954, Gonocephalum caesum Steven, 1829: 96. – Gebien 1910b: 322, 1939:
C. Jacot leg. 1 ex.; Vallée de Mt Orange, 1.500 m, R. 452; Ferrer 2000: 122.
Ellenberger leg. 1906, Basutoland: Machlatsa, 30.III. Hopatrum caesum (Steven, 1829). – Gemminger 1870: 1931.
1954, Gillarmed leg., 10 ex. (TMNH); Afrique du Sud:
Cape prov. “Gonocephalum capensis n. sp. Ardoin Type data. Syntypes: “Ex Insula Macarena (Ile
det.” [in litt.], 2 ex. (MNHN); OFS: Barberspan, 45 km Burbon) D. Roux”.
SW Lichtenburg, 29.V.1954, G. Rudebeck leg., 1 ex. Distribution. AFROTROPICAL REGION: Bourbon
(ZMUL); OFS Christiana N. Persson leg., 8 ex. (ZMUB); Isl., Mauritania.
256 D. IWAN, J. FERRER and M. RAŚ
Opatrum calcaripes Karsch, 1881: 59. – Champion 1895: 115. Hopatrum catenulatum Fairmaire, 1896: 19.
Gonocephalum calcaripes (Karsch, 1881). – Kolbe 1902: 578; Gebien Gonocephalum catenulatum (Fairmaire, 1896). – Gebien 1910b: 322,
1910b: 322, 1939: 452; Ferrer 1993a: 78, 1995a: 12, 2007: 92; Iwan 1939: 447; Kaszab 1952a: 463; Iwan and Löbl 2008: 263.
and Löbl 2008: 263.
Gonocephalum yelamosi Español et Viñolas, 1983: 37. – Ferrer
Type data. Syntypes (BMNH).“Kanara, Bombay,
1993a: 78, 1995a: 11, 2004: 84, 2007: 92 (syn.).
Andrewes leg.”.
Distribution. ORIENTAL REGION: India, Hima-
Type data. Opatrum calcaripes Karsch, 1881.
laya. PALAEARCTIC REGION (Iwan and Löbl 2008):
Holotype: male (ZMHB) “Rolas, Groeff (60988)”.
Gonocephalum yelamosi Español et Viñolas, Asia: India: Sikkim, Darjeeling District.
1983. Holotype: male (Mus. Barcelona), “Les Cases
d’Alcanar (Tarrangona), 14-III-1982 (T. Yélamos leg.)”
Paratypes: Granollers (X-1932); río Besós (XI-1940); G. celebense Kaszab, 1952
Móra d’Ebre (IV-1953); L’Ampolla (IX-1933); Les Cases Gonocephalum celebense Kaszab, 1952a: 558.
d’Alcanar (14-III-1982); Benicàssim (V-1956); El Grau
(IV-1957); Castelló de la Plana (IV-1946); Manises (III- Type data. Syntypes: 2 ex. (NHMB).
1959); Totana (V-1954); Pórtugos (VI-1959); San Roque Material studied. “Minahassa; Süd-Celebes; Cele-
(26-I-1956, III-1959); Cádiz (XII-1939, V-1940); Belén (26- bes, 1855 (leg. Pfeiffer)”.
VIII-1950); Nador (XII-1942); Ceuta (28-I-1930); Casa- Other depositories. ZMHG (2 ex.), NHMA (2 ex.).
blanca (V-1937); 31 males, 21 females (Mus. Barcelona). Distribution. ORIENTAL REGION: Indonesia (Ce-
Distribution. AFROTROPICAL REGION: Benin,
lebes).
Ghana, Guinea, Nigeria, Saint Thomas. PALAEARCTIC
REGION (Iwan and Löbl 2008): Europe: France (incl.
Corsica, Monaco); Portugal; Spain (incl. Gibraltar).
G. charbonnelae Ferrer, 2004
North Africa: Algeria; Morocco (incl. Western Sahara).
Gonocephalum charbonnelae Ferrer, 2004: 85.
PALAEARCTIC REGION (Iwan and Löbl 2008): Asia: Type data. Holotype (NHMB).
China: Hainan, Yunnan. Material studied. “Indus; India or. (coll. F. Bates)”.
Remarks. China (Kanton), India (Assam) (Rei- Other depository. BMNH (1 ex.).
chardt 1936). Distribution. PALAEARCTIC REGION (Iwan and
Löbl 2008): Asia: “Indus”.
Paratype: Namibie: Kavango prov., Rundu, I.1992, Gonocephalum contractum Gerstaecker, 1871: 61. – Gerstaecker
1873: 179; Gebien 1910b: 322, 1939: 451; Ferrer 1993a: 106, 1994:
Rautenstrauch leg., 1 female (ZSMB).
201, 2000b: 96, 2001: 230; Kaszab, 1971a: 227.
Distribution. AFROTROPICAL REGION: Angola,
Namibia.
Type data. Lectotype (designated by Ferrer 2000b:
96): male (ZMHB), “Lac Djipe, v. Decken (types
56624)”.
G. condamini Ardoin, 1963
Paratypes: Lac Djipe, v. Decken (types 56624), 4 ex.
Gonocephalum condamini Ardoin, 1963: 1379. – Ferrer 1995a: 15, (ZMHB).
2004: 84. Distribution. AFROTROPICAL REGION: Burundi,
Kenya, Malawi, Mozambique, Namibia (Caprivi Strep),
Type data. Holotype: male (MNHN), “Sénègal, Republic of South Africa (Basutholand), Ruanda, Tan-
Basse Cassamance, Fôret classé de Tobor, 13.XI.1961, zania, Zambia, Zimbabwe.
A. Villiers leg.”.
Distribution. AFROTROPICAL REGION: Benin,
Burkina Faso, Chad, Ghana, Nigeria, Senegal. G. controversum Gridelli, 1948
Type data. Holotype: female (BMNH), “Irak, Ama- Type data. Unknown.
ra, 18.II.1914, (under clods), W.E. Evans leg.”. Distribution. AUSTRALIAN AND PACIFIC RE-
Distribution. AFROTROPICAL REGION: Erithrea, GIONS: Torres Isl.
Ethiopia. ORIENTAL REGION: Burma, India. PALAE-
ARCTIC REGION (Iwan and Löbl 2008): Asia: Afgha-
nistan; India: Uttaranchal (Uttar Pradesh); Iran; Iraq; G. coriaceum Motschulsky, 1858
Pakistan; Saudi Arabia; Yemen (incl. Socotra).
Gonocephalum coriaceum Motschulsky, 1858 [1857]: 34. – Motschul-
sky 1866: 173; Harold 1876: 130; Marseul 1876: 96; Reitter 1904:
143; Gebien 1910b: 322, 1913: 6, 1939: 447; Schuster 1928: 989;
G. contractum Gerstaecker, 1871 Reichardt 1936: 112; Kaszab 1941a: 51, 1952a: 667, 1954b: 248;
Chûjô, 1966: 7; Kwon and Choi 1986: 106; Hua Li 2002: 138; Iwan
Opatrum contactum Gerstaecker, 1861: 61. – Champion 1895: 116. and Löbl 2008: 263.
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 259
Opatrum coriaceum (Motschulsky, 1858). – Lewis 1894: 381. G. cowardense Blackburn, 1894
Hopatrum coriaceum (Motschulsky, 1858) – Gemminger 1870:
1931. Gonocephalum cowardense Blackburn, 1894: 215. – Blackburn, 1907:
287; Gebien 1910b: 323, 1939: 450.
Type data. Unknown.
Distribution. PALAEARCTIC REGION (Iwan and Type data. Syntypes: “Cowards Springs” (BMNH).
Löbl 2008): Asia: China: Fujian (Fukien), Guandong Distribution. AUSTRALIAN AND PACIFIC RE-
(Kwantung), Guanxi (Kwangsi), Nei Mongol (Inner GIONS: South Australia.
Mongolia), Sichuan (Szechwan), Shandong (Shantung),
Zhejiang (Chekiang); Japan; Nepal; Russia: Far East;
South Korea; Taiwan (Formosa).
G. crassepunctatum Kaszab, 1952
Opatrum costatum Brullé, 1832: 218. Type data. Syntypes: 6 ex. (HNHM).
Hopatrum costatum Brullé, 1832. – Gemminger 1870: 1931. Material studied. “Trichinopoly; Fyzabad, United.
Gonocephalum costatum (Brullé, 1832). – Baudi 1875: 229; Seidlitz
Prov. (leg. R. W. G. Hingston); Tenasserim; Kowkereel
1894: 440; Reitter 1904: 143; Gebien 1910b: 322, 1939: 445; Schus-
ter 1928: 989; Reichardt 1936: 106; Kaszab 1939: 588, 1961a: 4,
(leg. Fea); Burma, Rangoon, VII.1935 (leg. L. N. Johri);
1967b: 551; Kühnelt 1965: 35; Grimm 1991: 28. 25. bis 30.XI.1934 (leg. R. Malaise); Hanoi”.
Gonocephalum costatum costatum (Brullé, 1832). – Ferrer 1993a: Other depository. BMNH (5 ex.).
87 (stat. subsp.), 1995a: 45; and Fattorini 2002: 160; Iwan and Löbl Distribution. ORIENTAL REGION: Burma, India,
2008: 263.
Vietnam. PALAEARCTIC REGION (Iwan and Löbl
Opatrum lineare Küster, 1849a: 54.
Hopatrum lineare Küster, 1849. – Gemminger 1870: 1932. 2008): Asia: Bhutan; Nepal.
Gonocephalum lineare (Küster, 1849). – Kraatz 1873: 438; Seidlitz
1894: 440; Reitter 1904: 143; Gebien 1910b: 322; Reichardt 1936:
106.
Gonocephalum curiosum Kaszab, 1952a: 460. – Schawaller 1997a: 5; Gonocephalum deliensis Kaszab, 1952a: 505. – Iwan and Löbl 2008:
Iwan and Löbl 2008: 263. 263.
Type data. Syntypes: 4 ex. (Frey Museum, coll. Type data. Holotype (HNHM).
Gebien, in München) NHMB. Material studied. “Deli; Nord-Ost-Sumatra”.
Material studied. “Bengal, Pusa (leg. H. L. An- Other depository. NHMB (1 ex.).
drewes); Bengalen, Raymahal; Sychel, 1863 (leg. Distribution. ORIENTAL REGION: Sumatra.
Bowring); Peneng, 1963 (leg. Bowring); Bihar, Saheb- PALAEARCTIC REGION (Iwan and Löbl 2008): Asia:
ganj (coll. Ind. Mus.); Calcutta; Assam; Tezpur, Mangal- China: Guanxi (Kwangsi).
dai Distr., 8–9.X.1910 (leg. Kemp); Burma; Katha Bur-
ma, Monnyin (leg. C. F. C. Beeson); Tenaserim”.
Other depositories. BMNH (11 ex.), HNHM (2 ex.), G. dentipes Kaszab, 1952
SDEI (2 ex.), ZMHG (1 ex.).
Gonocephalum dentipes Kaszab, 1952a: 655. – Hua Li 2002: 138;
Distribution. ORIENTAL REGION: Burma, India. Iwan and Löbl 2008: 263.
PALAEARCTIC REGION (Iwan and Löbl 2008): Asia:
Nepal. Type data. Syntypes: 11 ex. (NHMB).
Material studied. Kajo-Tanam (leg. Klein); Battak
Gebirge (leg. H. Fruhstorfer); Palaboen, 1870 (leg. J.
G. curlettii Ferrer, 2000 Xántus); Salak (leg. K. Kraepelin); Buitenzoorg, 1919
(leg. W. C. van Heurn); Lombok, Soembalon (leg. Ren-
Gonocephalum curlettii Ferrer, 2000b: 134.
sch); Sambalum (leg. H. Fruhstorfer); Sapit, 2000 ft
(leg. H. Fruhstorfer); Semongkat am Nordhang der
Type data. Holotype: female (MNHN), “Soudan
Batoe Langkat Gebirge (leg. Rensch); Sumbawa, Dom-
Egyptien, Roseires; Haut Nil Bleu, Ch. Alluaud leg.
poe (leg. Rensch).
1904, coll. Alluaud”.
Other depositories. BMNH (3 ex.), HNHM (2 ex.),
Distribution. AFROTROPICAL REGION: Sudan.
ZMHB (33 ex.), SDEI (11 ex.), ZMHG (3 ex.), ZSMG (3
ex.), MCSN (3 ex.).
Distribution. ORIENTAL REGION: Indonesia (Su-
G. curvicolle Reitter, 1889
matra, Java, Lesser Sunda Isl.). PALAEARCTIC RE-
Gonocephalum curvicolle Reitter, 1889: 705. – Champion 1895: 115; GION (Iwan and Löbl 2008): Asia: Taiwan (Formosa).
Reitter 1904: 142; Gebien 1910b: 323, 1939: 446; Schuster 1928: 989;
Gridelli 1934: 66; Reichardt 1936: 107; Kaszab 1952a: 654; Iwan and
Löbl 2008: 263. G. dentitibia airense Gridelli, 1948
Gonocephalum curvicollis Reitter, 1889. – Hua Li 2002: 138.
Gonocephalum airensis Gridelli, 1948: 28.
Type data. Unknown. Gonocephalum dentitibia airense Gridelli, 1948. – Ferrer 1993a: 92
(stat. subsp.), 1995a: 65, 2004: 88.
Distribution. PALAEARCTIC REGION (Iwan and
Löbl 2008): Asia: China: Nei Mongol (Inner Mongolia),
Type data. Holotype: male (MCSN), “Niger, Air,
Gansu (Kansu), Sichuan (Szechwan), Xizang (Tibet);
Agadez, «Tabello Air Est, 800 m. 24–26.VIII.1947, L.
India: Himachal Pradesh, Kashmir; Mongolia.
Chopard-Villiers leg.”.
Remarks. Mongolia (Hua Li 2002: 138).
Distribution. AFROTROPICAL REGION: Niger.
Gonocephalum dentitibia iwani Ferrer, 2010: 236. Gonocephalum dermestoides latum Ferrer, 1995: 54. – Ferrer 2004:
87.
Type data. Holotype: female (MIZ), “Livingstone,
N. Rhodesia, I.1942, W. Eichler leg.”. Type data. Holotype: male (ZSMB), “Kenya, env.
Distribution. AFROTROPICAL REGION: “North Malindi, 13–29.VIII.1983, H.J. Bremer leg.”
Rhodesia”. Paratypes: Kenya, env. Malindi, 13–29.VIII.1983,
H.J. Bremer leg., 22 ex. (ZSMB), 1 male (JFCS); Burun-
di: Bulumburu Boom, 10.VII.1985, Mühle leg., 1 female
G. demeteri Ferrer, 1995 (ZSMB).
Distribution. AFROTROPICAL REGION: Kenya,
Gonocephalrum demeteri Ferrer, 1995a: 30. Somalia.
G. devylderi Ferrer, 1995 South India, Mysore State, Shimoga Dist., Agumbe
Ghat, 2000 ft., V.1980, 1 male (J. Ferrer coll.).
Gonocephalum devylderi Ferrer, 1995a: 65.
Distribution. ORIENTAL REGION: India.
Chilka Lake, 24.X.1920 “at leight” (leg. Annandale); Somalie ex ital. Giuba, Belet Amin, VII.1934, Patrizi leg.
Orissa coast, nr. Puri, Sur Lake, 19.VIII.1911 (leg. (Type!) (HNHM);
Annandale and Gravely); Anuradhapura (Low coun- Distribution. AFROTROPICAL REGION: Somalia,
try), 8–20.X.1911 (leg. N. Annandale); Jissa, 29.VII. Zimbabwe.
1921, 3.VIII.1921 (leg. S. P.)”.
Distribution. ORIENTAL REGION: India, Sri
Lanka. G. feae Gebien, 1920
Gonocephalum foveoseriatum Chatanay, 1914: 470. – Gebien 1939: Gonocephalum fuscosetosum Chatanay, 1917: 250. – Gebien 1939:
451; Ferrer 1993a: 99, 1996a: 61, 2000b: 119. 448; Kaszab 1952a: 612.
Type data. Syntype: 1 male (MNHN), “Kenya: Ch. Type data. Syntypes: “Indochina: Cochinchina,
Alluaud et Jeannel leg. oct. 1911, St. Côté d’Afrique or. coll. Chatanay”, 1 male, 1 female (MNHN).
anglaise, Ile de Mombasa”. Distribution. ORIENTAL REGION: Vietnam.
Distribution. AFROTROPICAL REGION: Kenya.
Type data. Holotype: male (TMNH), (Burkina Type data. Holotype (NHRS): “Oman, prés de
Faso), “Ob. Volta, Pundu, Olsufiew leg.” Quarayar, 100 m h, aux lumières, (in footchills) 12.III.
Paratypes: Ob. Volta, Pundu, Olsufiew leg., 1 female 1990, M. D. Gallagher leg. ref. n. 8243”.
(TMNH); 1 male (id., JFCS). Paratypes: Oman: Baushar, 140 m h, 3.IV.1991
Distribution. AFROTROPICAL REGION: Burkina (JFCS); Oman: Siya, M. D. Gallagher leg. ref. n. 5410,
Faso. 21.III.1991, Natural History Museum, Oman.
Distribution. PALAEARCTIC REGION (Iwan and
Löbl 2008): Asia: Oman.
G. freudei Kaszab, 1960
G. gebienianum Kaszab, 1952 Nelspruit, 26.II.1986 no. 2427, T.W. Drinkwater leg., 6
ex. (MZUF); Vryburg, 14.III.1985, id. no. 1767, id., 1 ex.
Gonocephalum Gebienianum Kaszab, 1952a: 470. – Kaszab 1979a:
(MZUF); Beaufort, Tiddbury, I,1961 (dans l’humus) N.
260.
Gonocephalum gebienianum Kaszab, 1952a. – Iwan and Löbl 2008: Leleup leg., 16 ex. (TMNH); Natal (ex coll. Oberthür), 1
263. ex. (MNHN); Tugela river, near Kranskop, Lawrence,
Hacke, leg., 2 ex. (JFCS); Natal\ “Opatrum lateritium
Type data. Syntypes: 29 ex. (SDEI). Buq.coll. Thomson”, 1 ex. (IRSN); Natal, 1901, H. Ertl
Material studied. “Darjiling Distr., 1–3000 ft, V.1912 leg., 2 ex. (TMNH); id. 1904, Pauks leg., 16 ex. (TMNH);
(coll. Lord Carmichael); Darjeeling, Gopaldhara, BW., Zulu(land), 22.VI.(1904?), I. Träghård leg., 1 ex.; id.
4720 ft, 1914 (leg. H. Stevens); Malang; Preanger (leg. 9.I.–7.II.(1904?), 1 ex. (NHRS); Durban, id. 16.XI–I.1904,
P. F. Sijthoff); Pengalengan, 4000 ft (leg. H. Fruhstor- id., 11 ex. (NHRS); Isipingo, A.H.N., 1 ex. (JFCS); Isipin-
fer); Lombok, Soembalon (leg. Rensch)”. go, A.H.N., 28.XI.1958, L. Schulze leg., 35 ex. (TMNH);
Other depositories. BMNH (3 ex.), NHMB (2 ex.), Kloof, 1.500 fts, IX.1926, R.E. Turner leg., 6 ex.
ZMHB (3 ex.). (TMNH); Tongaat, VII.1950, Miss. R. Hoffmann leg., 1
Distribution. ORIENTAL REGION: India, Indone- ex. (JFCS); Tongaat, V.1953, mrs. Koch leg., 1 ex.
sia (Java, Lesser Sunda Isl.). PALAEARCTIC REGION (TMNH); Port Natal, J. Schulze leg.\” Gonocephalum
(Iwan and Löbl 2008): Asia: India: Sikkim (Darjeeling bettoni Blair in litt.”, 1 ex. (BMNH); Empangeni Uni-
District). versity, IX.–X.1975, P.E. Reavel leg., 1 ex. (TMNH);
Orange Free State: Bloemfontein no. 2059, T.W.
Drinkwater leg., 3 ex.; Brandfort, 26.III.1985, no. 2088,
G. geneirotundum Ren, 1998 3 ex.; Globbelaarl, 5.III.1986, id. no. 2536, 1 ex.; Wes-
selsbron, 31.I.1979; no.1162, id. 10 ex.; Winburg,
Gonocephalum geneirotundum Ren, 1998: 111. – Iwan and Löbl
2008: 263. 9.IV.1981, no.1453, id., 3 ex.; 16.IV.1985, no.2116, id.,
1 ex.; Wolmaransstad, 30.III.1982, no.1496, id., 1 ex.
Type data. Unknown. (MZUF); Mozambique: Umbeluzi sidt. 6.XII.1952, C.
Distribution. PALAEARCTIC REGION (Iwan and Koch leg., 1 ex. (JFCS); env. Vila Péry; bord du
Löbl 2008): Asia: China: Zhejiang (Chekiang). M’zingazé, Pied de Cabeza de Velho, 20.V.1929, P.
Lesne leg., 1 ex.; Sentier de Chizombeze, Vila Péry,
XI.1929, id., 2 ex. (MNHN); Lourenço Marques,
G. germari germari Ferrer, 2000 6.II.1950, P. Valerio leg., 148 ex.; id. VIII.1950, id., 61 ex.
(TMNH); Pafuri, IV.1981, D.W. Rorke leg., 14 ex.
Gonocephalum germari germari Ferrer, 2000b: 116. – Ferrer 2004: 90. (TMNH); Zimbabwé: “Rhodesia”\”G. rhodesianum n.
sp. Blair det.” (in litt.), 1 ex. (BMNH); Rusape, 25.XII.
Type data. Holotype: male (TMNH), “Afrique 1992, K.P. Adlbauer leg., 1 ex. (BMNH); Chianigani,
du Sud: Transvaal, Ohrigstad, 15 km S, 11.XI.1980, 27.XI.1992, G. Wallsberger leg., 1 ex. (TMNH); Pen-
S. Endrödy-Younga leg.”. tridge, 11.I.1928, R.H.R. Stevenson leg., 1 ex. (TMNH);
Paratypes: Afrique du Sud: Transvaal, Ohrigstad, Zimbabwe, VIII.1930, G. von Son leg., 2 ex. (TMNH).
15 km S, 11.XI.1980, S. Endrödy-Younga leg., (HNHM); Distribution. AFROTROPICAL REGION: Mozam-
Ohristad, 1.VII.1962, N. Leleup leg., 21 ex. (TMNH); bique, Republic of South Africa (Transvaal, Natal, Cape
“Cape prov. Krtz coll” (ex coll. Kraatz) Haag- Province), Zimbabwe.
Rutenberg, 3 ex. (SSM); Cap B Spei, Dunbrody\”Opa-
trum micans Germ. det. L. Fairmaire, 1 ex. (IRSN);
Shilovane, 1906-1907, N.A. Junod leg., 3 ex. (MNHN); G. germari hemmingi Ferrer, 2000
Kalahari, Kabula Bula, Chobe River, 11–24.VII.1930,
Vernay Transvaal Mus. exp. (sp. dubiosa, gra- Gonocephalum germari hemmingi Ferrer, 2000b: 118. – Ferrer
nohispidum? Det. H.J. Hesse, 1 ex. (TMNH); S. Afr. 2004: 90.
Barbeton, X.1927, G.V.D. leg. granulipennis mihi s.
sp. aut spec. nov. det. Gridelli 1950, ex coll. Koch, 1 ex. Type data. Holotype: male (TMNH), “Somalie:
(TMNH); Barbeton, Miss. L. C de Beer leg. 1900, 6 ex. North Region, Talch 10 (2.000 fts) IX–XI.1960, C.F.
(MNHN); Hluluwe Game Reserve, 18.IV.1951, no. 277; Hemming leg.”.
Brinck, Rudebeck leg., 1 ex. (TMNH); Margate, I.1943, Paratypes: Somalie: North Region, Talch 10 (2.000
W.G. Kobrow leg., 2 ex. (TMNH); Victoria East, fts) IX–XI.1960, C.F. Hemming leg., 51 ex., 5 ex. (JFCS);
VII.1948, M. Mc Gilliway leg., 1 ex. (TMNH); Pretoria, Galgalo, IX.1950, C. Koch leg., 49 ex. (TMNH); Galgalo
X.1958, L. Schulze leg., 26 ex. (TMNH); Pretoria, Oasis, 10.X.1973, Spediz. Boilog. in Somalia (S.B.S),
II.1962, L. Vari leg., 44 ex. (TMNH); Branvlei, VIII.1937, 5 ex., (MZUF 8401), 1 ex. (JFCS); Afgoi, 29.IX.1973,
1 ex. (TMNH); Port Shepstone, I.1953, 13 ex. (TMNH); S.B.S., 4 ex. (MZUF 8402).
266 D. IWAN, J. FERRER and M. RAŚ
Gonocephalum gestroi mali Ferrer, 1995: 34. Penthicus gracilis Bates, 1879: 482. – Bates 1890: 19.
Loboderus gracile Bates, 1879: 482. – Bates 1890: 75.
Lobodera gracile (Bates, 1879). – Reichardt 1936: 169.
Type data. Holotype: male (IRSN), “R. Cen-
Gonocephalum gracile (Bates, 1879). – Gebien 1939: 447; Kaszab
treafrique, Bangui, IV.1981, N. Degallier leg.” 1952a: 546, 1961b: 351, 1977: 257, 1978b: 230; Schawaller and
Paratypes: R. Centreafrique, Bangui, IV.1981, N. De- Skopin 1978: 230; Schawaller 1997a: 6; Hua Li 2002: 138; Iwan and
gallier leg., 1 female (IRSN); Rep. Centreafrique, Ouban- Löbl 2008: 263.
Gonocephalum ruficorne Gravely, 1915: 250. – Kaszab 1952a:
gui Chari, Bangui, 1.III.1968, ex. coll. Breuning, 3 ex.
676.
(MRAC); Cameroun: Neu Quella, IV.1913, S.G. Houy leg., Gonocephalum curvicolle Gridelli, 1934: 66. – Kaszab 1952a: 676;
1 ex. (BMNH); Neu (2475-95) Tessman leg., 1 ex. (ZMHB); hom. nov. nom. preoccup., nec Gonocephalum curvicolle Reit-
Cote d’Ivoire: Bouake, 1946, R. Delattre leg., 1 male, ter, 1889: 705.
1 female coll. Noël Mal, Bruxelles; Tchad: Dar Band mèr.
Krebedje, Fort Sibuti, IX.1904 Dr Decorse leg., 2 ex. Type data. Unknown.
(MNHN); Zaire: Njam, Schlüter leg., 1 female (ZMHB). Distribution. PALAEARCTIC REGION (Iwan and
Distribution. AFROTROPICAL REGION: Came- Löbl 2008): Asia: China: Xizang (Tibet), Xinjiang (Sink-
roon, Central African Republic, Chad, Ivory Coast, iang); India: Himachal Pradesh, Kashmir, Sikkim (Dar-
Zaire. jeeling District); Nepal.
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 267
G. granulatum nigrum (Küster, 1849) Hopatrum pruinosum Fischer von Waldheim, 1844. – Gemminger
1870: 1933.
Opatrum nigrum Küster, 1849a: 58.. Gonocephalum pruinosum (Fischer von Waldheim, 1844). – Schus-
Hopatrum nigrum Küster, 1849. – Gemminger 1870: 1932. ter 1934: 128.
Gonocephalum nigrum (Küster, 1849). – Sturm 1843: 152; Mulsant Opatrum ragusanum Küster, 1849b: 52.
1854: 168; Brisout 1863: 45; Baudi 1875: 702; 1875: 231; Tenebrion- Hopatrum ragusanum Küster, 1849. – Gemminger 1870: 1933;
iti, p. 67; Rey 1892: 42; Seidlitz 1894: 446 (= Opatrum pusillum Gonocephalum ragusanum (Küster, 1849). – Seidlitz 1894: 446; Reit-
Fabricius, 1791); Reitter 1904: 144 (= Opatrum pusillum ter 1904: 144; Gebien 1910b: 325.
Fabricius, 1791); Gebien 1910b: 325 (= Opatrum pusillum Gonocephalum pusillum var. subnodosum Rey, 1892: 42.
Fabricius, 1791), 1939: 444 (= Opatrum pusillum Fabricius, Gonocephalum subnodosum Rey, 1892. – Gebien 1910b: 325
1791); Reichardt 1936: 102 (= Opatrum pusillum Fabricius, (= Gonocephalum pygmaeum Steven, 1829), 1939: 443
1791). (= Gonocephalum pygmaeum Steven, 1829).
Gonocephalum granulatum nigrum (Küster, 1849). – Ferrer 1993a: Opatrum viennense Duftschmid, 1812: 293.
86 (stat. subsp.), 1995a: 42; Leo and Fattorini 2002: 160; Iwan and Gonocephalum viennense (Duftschmid, 1812). – Küster 1849b: 53;
Löbl 2008: 263. Redtenbacher 1849: 597, 1858: 569, 1874: 99; Bach 1856: 216; Rey
1892: 42; Seidlitz 1894: 446; Desbrochers 1901: 93; Reitter 1904:
144; Gebien 1910b: 325.
Type data. “Im südlichen Frankreich bei Montpel-
Hopatrum viennense (Duftschmid, 1812). – Gemminger 1870: 1934.
lier. Männchen und Weibchen zur Beschreibung aus
Sturms Sammlung”.
Type data. Opatrum pusillum Fabricius, 1791.
Distribution. PALAEARCTIC REGION (Iwan and
Lectotype (designated by Ferrer 1995a: 43) Syntypes:
Löbl 2008): Europe: France (incl. Corsica, Monaco);
1 male, 4 female, «pusillum» sans autre indication, le
Italy (incl. Sardinia, Sicily, San Marino); Spain (incl.
mâle en tres mauvais état, sans avant corps, désigné ici
Gibraltar).
comme Lectotype (ZMC);
Remarks. Palearctic Region: Turquie, Bulgarie,
Opatrum interstitiale Küster, 1849. “In Montene-
Yougoslavie, Greece (Leo and Fattorini 2002).
gro bei Cettigne auf trockenen Grasplätzen”.
Opatrum nitidulum Küster, 1849. “In der Türkey,
G. granulatum pusillum (Fabricius, 1791) von M. Wagner gesammelt”.
Opatrum ragusanum Küster, 1849. “Bei Ragusa
Opatrum pusillum Fabricius, 1791: 91. – Herbst 1793: 224; Fabricius ziemlich selten an trocken Abhängen”.
1801: 91; Sturm 1807: 172; Duftschmitd 1812: 293; Thunberg 1821: Distribution. PALAEARCTIC REGION (Iwan and
54; Besser 1832: 17; Krynicki 1832: 135; Bach 1856: 216; Redten-
Löbl 2008): Europe: Azerbaijan; Albania; Armenia;
bacher 1858: 569, 1874; Baudi 1876: 67; Lindemann 1888: 58; Sei-
dlitz 1891: 551, Muller 1920: 200; Portvein 1934: 14; Ogloblin and
Austria; Bulgaria; Croatia; Czech Republic; Hungary;
Kolobowa 1927: 19. Greece (incl. Crete); Italy (incl. Sardinia, Sicily, San
Hopatrum pusillum Fabricius, 1791. – Gemminger 1870: 1933. Marino); Macedonia; Poland; Romania; Slovakia; Spain
Gonocephalum pusillum (Fabricius, 1791). – Brisout 1863: 45; Rey (incl. Gibraltar); Russia: South European Territory;
1892: 42; Seidlitz 1894: 446; Reitter 1904: 144, 1911: 335; Gebien
Ukraine; Yugoslavia (Serbia, Montenegro). North
1910b: 324, 1939: 444; Schuster 1928: 989; Reichardt 1936: 102;
Kaszab 1939: 589, 1940: 6, 1959:, 1961a: 4, 1961c: 173, 1967: 333, Africa: Morocco (incl. Western Sahara). Asia: Afghan-
1967b: 550; Español 1959: 246, 1969: 85; Samedov 1963:; Kühnelt istan; China: Ningxia (Ningsia), Xizang (Tibet); Iran;
1965: 35; Canzoneri 1968: 103, 1970: 75; Skopin 1968:, Marcuzzi Kazakhstan; Mongolia; Syria; Turkmenistan; Turkey.
1970: 89; Knor 1973: 99; Medvedev 1974, 1990: 1701; Minoranskij
Remarks. Algieria (Gemminger 1870: 1933).
1976; Angelov and Medvedev 1981: 307; Kaluzhnaya 1982: 74;
Grimm 1985: 17; Canzoneri and Vienna 1987: 19; Medvedev and
Küster (1849b: 53) and Redtenbacher (1849: 597,
Lobanov 1990: 187; Schawaller 1996: 9. 1858: 569, 1874: 99) listed species Gonocephalum
Gonocephalum granulatum pusillum Fabricius, 1791: 91. – Ferrer viennense Duftschmid, 1812 which are interpreted by
1993a: 86 (stat. subsp.), 1995a: 43; Iwan and Löbl 2008: 263. Ferrer (2009: 123) as Gonocephalum pygmaeum
Opatrum interstitiale Küster, 1849a: 59.
(Steven, 1829).
Hopatrum interstitiale Küster, 1849. – Gemminger 1870: 1932.
Gonocephalum interstitiale (Küster, 1849). – Seidlitz 1894: 446;
Reitter 1904: 144; Gebien 1910b: 325.
Opatrum muricatum Brullé, 1832: 215. G. greensladei Kaszab, 1980
Hopatrum muricatum Brullé, 1832. – Gemminger 1870: 1932.
Gonocephalum muricatum (Brullé, 1832). – Seidlitz 1894: 446; Reit- Gonocephalum greensladei Kaszab, 1980: 34.
ter 1904: 144; Gebien 1910b: 325, 1939: 444.
Opatrum nitidulum Küster, 1849a: 56.
Hopatrum nitidulum Küster, 1849. – Gemminger 1870: 1932.
Type data. Holotype: male (HNHM), “Guadalca-
Gonocephalum nitidulum (Küster, 1849). – Seidlitz 1894: 446; Reit- nal: Kukum, 29. Okt. 1963, P. G. (No 9734, im Mus.
ter 1904: 144; Gebien 1910b: 325. Budapest)”.
Opatrum parvulum Lucas, 1846: 334. Paratypes: Guadalcanal: Mt. Austin, 15. Jan. 1963,
Hopatrum parvulum Lucas, 1846. – Gemminger 1870: 1933.
P. G. (No 3444), 1 ex (HNHM); Vitale, 6.Jan.1963, P. G.
Gonocephalum parvulum (Lucas, 1846). – Seidlitz 1894: 440
(= Opatrum prolixum Erichson, 1843). (No 3309), 4 ex. (HNHM); Guadalcanal: Vitale, Apr.
Opatrum pruinosum Fischer von Waldheim, 1844: 128. 1965, P. G. (16798), 2 ex. (HNHM); Guadalcanal: Dez.
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 269
1920, Jan. 1921, Feb. 1921, J. A. Kusche, 3 ex. (DECB); Type data. Holotype: male (MCSN), “Ethiopie
Guadalcanal: Honiara, 13.–16. Nov. 1953, J. D. B., 1 ex. méridionale: Gota. VIII.1936, Lomi leg. (“Typus,
(BMNH); Tulagi: 10. März 1936, R. A. Lever, 2 ex. G. lomii n. sp. Gridelli det.”) (in litt.) Mojo Atschavo,
(BMNH). 2.VI.1900, 1 ex.”.
Distribution. AUSTRALIAN AND PACIFIC RE- Paratypes: Harar, 14.V.1900 (12); 5.III.1900, 1 ex.
GIONS: Guadalcanal. (ZMHB); Bisidimo, Hararge prov. 1.500 m, XI.1983–
I.1984, Verena Meyer leg., 3 ex. (ZSMB); Somalie:
Hargeisa, 1.300 m, 21–31.VII.1980, Bremer leg., 9 ex.
G. gridellii Koch, 1953 (ZSMB); Tanzanie: Tanganyka, 2 liles to Lake Man-
yara, SE shore, 3.150 fts. 1–26.VI.1965, J. Szunyoghi
Gonocephalum gridellii Koch, 1953: 288. – Ferrer 1993a: 93, 1995a:
leg., 7 ex. (HNHM).
66.
Distribution. AFROTROPICAL REGION: Djibouti,
Ethiopia, Tanzania.
Type data. Paratype: G. n. sp. Det. Gridelli 1951,
male.
Distribution. AFROTROPICAL REGION: Republic
G. grimmi Ferrer, 1995
of South Africa (Transvaal).
Gonocephalum grimmi Ferrer, 1995a: 52.
G. gridellianum Kaszab, 1952 Type data. Holotype: male (ZMB), “Nova Rodondo,
Angola”.
Gonocephalum Gridellianum Kaszab, 1952a: 537.
Paratype: Nova Rodondo, Angola, 1 female (ZMB).
Distribution. AFROTROPICAL REGION: Angola,
Type data. Syntypes: 3 ex. (MCSN) “Nongkodjad-
Republic of Congo.
jar, 4000 ft (leg. J. P. A. Kalis)”.
Distribution . ORIENTAL REGION: Indonesia
(Java). G. gruveli Ferrer, 2004
Gonocephalum guignoti Kaszab, 1960a. – Kaszab 1968b: 20, 1970c: Opatrum hadroides Wollaston, 1851) (error: not “hadroide”);
148, 1973b: 55; Iwan and Löbl 2008: 264. Hua Li 2002: 139 (= Gonocephalum reichardti Gebien, 1939).
Gonocephalum Reichardti Gebien, 1939: 447 (replaced name).
– Kaszab 1952a: 584; Iwan and Löbl 2007: 735 (syn.).
Type data. Holotype, Allotype (NHMB).
Material studied. “Afghanistan: Umgebung von
Kabul, Tangi Gharuh am Kabulfluß, 40 km südöstlich Type data. Syntypes: probably in MNHN.
von Kabul, 1600 m, 10.V., 8.VII., 20.IX.1952, J. Klap- Distribution. PALAEARCTIC REGION (Iwan and
perich; Panchirtal, Bazarak, 2200 m, 27.VI.1952, Löbl 2008): Asia: China: Beijing (Peking or Peiping),
J. Klapperich; Salangtal, Aghelekan, 1900 m, 12.X.1952, Sichuan (Szechwan).
J. Klapperich; Salangtal, Ejan, 2050 m, 11.X.1952,
J. Klapperich; Nuristan, Bashgultal, 1100 m, 14.VII.
1953; 9.XI.1953, J. Klapperich”. G. hauschildi Kaszab, 1952
Other depositories. 71 paratypes (NHMB). Gonocephalum Hauschildi Kaszab, 1952a: 498. – Kaszab 1965a: 113.
Distribution. ORIENTAL REGION: India.. PALAE- Gonocephalum hauschildi Kaszab, 1952a. – Schawaller 1997a: 6;
ARCTIC REGION (Iwan and Löbl 2008): Asia: Afghan- Iwan and Löbl 2008: 264.
istan; Pakistan.
Type data. Syntypes: 2 ex. (HNHM).
Material studied. “Darjeeling; Sikkim, Namsov, XI.
G. gustafssoni Ferrer, 1995 1920 (leg. H. Stevens); Bengal, E. Duars (leg. H.
Stevens); Kaligad, Dehra Dum, 19.VI.1921 (leg. Dr.
Gonocephalum gustafssoni Ferrer 1995a: 73.
Cameroon); Sitapahar R., Chittagong H. T., 11.I.1925
(leg. C. F. C. Beeson)”.
Type data. Holotype: male (NHRS), “India (sans
Other depositories. BMNH (9 ex.), NHMB (2 ex.),
autres indications)”.
ZSMG (2 ex.).
Paratypes: N. India, Rishikesh (30.07°N, 78.19°E),
Distribution. ORIENTAL REGION: India, Thai-
VI.1986, K. Werner leg., female (ZSMB); India (sans
land, Vietnam. PALAEARCTIC REGION (Iwan and Löbl
autres indications) 1 male 2 females.
2008): Asia: China: Guanxi (Kwangsi); India: Sikkim
Distribution. PALEARCTIC REGION: India: Rishi-
(Darjeeling District), Uttaranchal (Uttar Pradesh)
kesh, Thar (Ferrer 1995a).
“Thibet”; Nepal.
G. himalayense Kaszab, 1952 Gonocephalum hiranoi Akita et Masomoto, 2003b: 173. – Iwan and
Löbl 2008: 264.
Gonocephalum himalayense Kaszab, 1952a: 528. – Schawaller
1997a: 8; Iwan and Löbl 2008: 264. Type data. Holotype: 1 ex., “Nishihama~Yana-
gishima-kaigan, Chigasaki City, Kanagawa Pref., Hon-
Type data. Holotype (HNHM). shu, Japan, 23.III.1998, Y. Hirano leg.”
Material studied. “Kurseong, Verschraeghen, 1904 Paratypes: Nishihama~Yanagishima-kaigan, Chi-
(coll. Baron Moffaerts); Sikkim; Calcutta, 23.IV.1910, gasaki City, Kanagawa Pref., Honshu, Japan, 23.III.
“under brichs by rubbin heap” (coll. Ind. Mus.)”. 1998, Y. Hirano leg., 2 ex.; Banyu, Hiratsuka City, Ka-
Other depository. BMNH (2 ex.). nagawa Pref., 15.III.1992, Y. Hirano leg, 5 ex.
Distribution. ORIENTAL REGION: India. PALAE- Distribution. PALAEARCTIC REGION (Iwan and
ARCTIC REGION (Iwan and Löbl 2008): Asia: Bhutan, Löbl 2008): Asia: Japan (Honshu).
Nepal, India: Sikkim (Darjeeling District).
Type data. Holotype: 1 male (MCSN), “Cameroun Paratypes: Tanzanie, Africa Or. Katona leg. 1904-
Duli”. 1905, 2 females (ZSMB); Sansibar, Kitui, Hildebrandt
Distribution. AFROTROPICAL REGION: Burkina leg., 1 male, «D.O.A.»: Tanga, O. Kickel and F. Fischer
Faso, Cameroon, Central African Rrepublic, Chad, leg., 1 female; Pangani steppe: Membo Massinde,
Eritrea, Republic of Congo, Zair. 1 female; Magamba Massinde (700–1600 m. h.), I.1906,
ex. Coll. Dr. C. Schröder, 1 male; Rabai: I–II.1929, A.F.J.
Gedye leg., 1 female (dèterminès par Blair comme des
G. inaequale danielssoni Ferrer, 1995 G. exiguum Blair in litt.) (BMNH); Nioa, IV.1915, Math-
ner leg., 1 ex. [museum]; Kikogwe id., 1 ex.; Ramisi,
Gonocephalum inaequale danielssoni Ferrer, 1995a: 28. – Grimm XI.1915, id., 1 ex.; Trockenwald-Mtotoliou, IV.1916,
2002: 368. «G. prolianum Gebien det.», 2 ex. (ZMHB); Kikogwe,
id., 1 ex. (MCSN).
Type data. Holotype: male (ZMUL), “Gambia, Kotu Distribution. AFROTROPICAL REGION: Tanzania.
stream près de Bakau (3 km SW), 22–23.XI.1977, Remark. Español and Viñolas (1983: 36) deter-
R. Danielsson leg.”. mined specimens belonging to Gonocephalum indic-
Paratypes: Gambia, Gunjur (5 km SW), 13.XI.1977, tum indictum Ferrer (described in 1995) as Gonoce-
R. Danielsson leg., 1 female (ZMUL); Kotu stream phalum angusticolle (Gerstaecker).
14.XI.1981, J.A. Janzon leg., 1 male (RSM); Sénègal:
Bambay, 1945, Ribec leg., 2 females, 1 male (MNHN);
Mpak, 11 km S Ziguinchor, 8.XI.1977, R. Danielsson et G. indictum longimanum Ferrer, 1995
al., 2 ex. (ZMUL); «Sénègal» «angustatum Dejean»
ex coll. Thompson, 1 ex. (IRSN); Guinea, Deyrolle «an- Gonocephalum indictum longimanum Ferrer, 1995a: 21. – Ferrer
1995b: 40, 1996a: 60, 1998a: 40.
gustatum», 1 ex. (ZMHB).
Distribution. AFROTROPICAL REGION: Gambia,
Type data. Holotype: male (ZSMB), “Kenya, envi-
Guinea, Senegal.
rons de Malindi, 13–29.VIII.1983, leg. H.J. Bremer (sub
lapidae)”.
Paratypes: Kenya, Taru Mombasa, F. Thomas leg.,
G. inaequale inermipes Gridelli, 1945 1 female (BMNH); Kenya: Malindi, 10.XI.–6.XII.1989,
Gonocephalum inermipes Gridelli, 1945: 26.
L. Bartolozzi leg. (Miss. Accad. Naz. Lincei), 2 females
Gonocephalum inaequale inermipes Gridelli, 1945. – Ferrer 1993a: (7815 MZUF), 1 male (JFCS); Malindi, 27.IX–14.X.1992,
73 (stat. subspec.), 1995a: 28 [“inermiceps” lapsus]. L. Bartolozzi leg., 1 female (MZUF 8576); Somalie: Gio-
har, 10/18?–28.IV.1968 (Sped. Biologica in Somalia,
Type data. Syntype: 1 male (ZSMB), “Abyssinie, S.B.S.), 1 female (MZUF PT 892).
Raffray». Semienie 719 saml. Haag Rutenberg, («elon- Distribution. AFROTROPICAL REGION: Kenya,
gatum Guér.»)”. Somalia.
Distribution. AFROTROPICAL REGION: Burkina
Faso, Chad, Ethiopie.
G. indicum (Kaszab, 1952)
Gonocephalum inquinatum inquinatum (Sahlberg, 1923). – Ferrer Paratypes: Congo, Luc, I.F.A.N. (ex coll. Fleutiaux),
1993a: 98, 2000b: 97, 2004: 89; Grimm 2002: 369. 1 male, 1 female (MCSN); R. P. Congo: Loudima,
Gonocephalum pubescens var. inquinatum Sahlberg, 1923. – Gebi-
en 1939: 451.
XII.1963, Balogh et Csiki leg., 4 ex. (HNHM); Cameroun:
Victoria, “simplex det. Staudinger” \”suberinitum
mihi in litt. det. Gridelli, 1937”, 1 female (MCSN); R. D.
Type data. Opatrum inquinatum Sahlberg, 1923.
Congo: Ngowa Kwango, 7.X.1937, R.P. Martens leg.
Lectotype (designated by Ferrer 2000b: 97): male
(inquinatum sahlbergi Ferrer in litt.), 27 ex. (IRSN);
(NHRS), “Sierra Leone, Afzelius leg. O. inquinatum,
id. XII.1938, 24 ex.; Kimbouko River, Kinsamba,
types”.
Bambesa, III.1937, J.M. Vrigdagh leg., 11 ex.; Libenge,
Paralectotype: Sierra Leone, Afzelius leg. O. inqui-
Miss, Mawuya, 1947, R. Cremer et M. Neuman leg., 27
natum, types, 1 female (NHRS).
ex.; Kiwit Kiriati, 7.VI.1981, P. Lesire leg., 19 ex. (ex
Distribution. AFROTROPICAL REGION: Burkina
coll. Noël Mal) (IRSN); Haute Congo, Bambili, 1907,
Faso, Cameroon, Chad, Gambia, Ghana, Guinea, Gui-
Ribotti leg., 1 ex. (MCSG); Moyen Congo: Forêt La
nea Bissau, Ivory Coast, Niger, Nigeria, Republic of
Mayombée, 1937, Aubert de la Rue leg., 1 ex. (MNHN);
Congo (Bazzaville-Congo), Senegal, Sierra Leone,
R. Centrafrique; Bangui, V.1981, N. Degallier leg.,
Togo.
13 ex. (ex coll. Noël Mal) (IRSN); Forêt la Maboqué,
IV.1964, J. Carayon leg. (G. inquinatum sahlbergi
Ferrer in litt.) (MNHN); Bukoko, V.1904, Pujol leg., 3 ex.
G. inquinatum palisoti Ferrer, 2000
(MNHN).
Gonocephalum inquinatum palisoti Ferrer, 2000b: 100. Distribution. AFROTROPICAL REGION: Came-
roon, Central African Republic, Republic of Congo.
Type data. Holotype: male (TMNH), “Angola, SW
Angola, Ongueria, VI.1954, J. Balfour leg. Vernay exp.”.
Paratypes: Angola, SW Angola, Ongueria, VI.1954, G. insulanum (Olliff, 1887)
J. Balfour leg. Vernay exp., 57 ex. (TMNH); 35 ex. id. Opatrum insulanum Olliff, 1887: 1006. – Champion 1895: 116.
(BMNH); 10 ex. (JFCS); Vana Lombe, VI.1954, Vernay Gonocephalum insulanum (Olliff, 1887). – Gebien 1910b: 323, 1939:
Exp., 4 ex. (TMNH); Dundo, II.1949, A. Barrios Macha- 449; Kaszab 1952a: 643, 1985a: 26.
do leg., 2 ex. (TMNH); Dundundo (Dundo) Forêt de gal-
lerie, 16.II.1954, id., 2 ex. (TMNH); Negola, 13.V.1974, Type data. Syntype: “Norfolk Isl.”.
W.D. Haache leg., 2 ex. (TMNH). Distribution. AUSTRALIAN AND PACIFIC RE-
Distribution. AFROTROPICAL REGION: Angola. GIONS: Norfolk Isl.
Gonocephalum parcepilosum Gridelli, 1940 (1939): 242. Gonocephalum interruptum Gridelli, 1948: 31. – Ferrer 1993a: 100,
Gonocephalum inquinatum parcepilosum Gridelli, 1940 – Kaszab 2000b: 103.
1963a: 345; Ferrer 1993a: 98, 2000b: 98, 2004: 90; Lillig and Bremer
2002: 69. Type data. Syntypes: female (MCSN), “B.O.A.; lit-
torale Chat. = interruptum Blair in litt. det. Gridelli”;
Type data. Holotype: male (MCSN), “Tessenei, type female (BMNH), “B.O.A. Upangwa, G. inter-
Antonio Remedeli”. ruptum n. sp. Blair det.”; type female (HNHM),
Remark. Gridelli designated only one specimen as “B.O.A., Upangwa” (in coll. Kaszab); Abyssinie, coll.
type (Holotype), but description of the subspecies Kaszab, 1 male (HNHM); Upangwa, XII.1912, 1 male
included specimens listed in the studied material: (ZMHB).
(MZUF) Adi Ugri, Andreini leg., luglio 1902. Distribution. AFROTROPICAL REGION: Ethiopia,
Distribution. AFROTROPICAL REGION: Chad, Somalia, Upangwa.
Eritrea, Senegal, Sudan.
G. japanum Motschulsky, 1860 A. Voehzkov leg. \ “G. juanensis n. sp. Blair det.”
in litt.”
Gonocephalum japanum Motschulsky, 1860: 16. – Motschulsky 1866:
Paratypes: Mozambique: San Juan de Nova, Canal V,
173; Harold 1878: 77; Gebien 1910b: 323, 1939: 447; Schuster 1928:
990; Reichardt 1936: 111; Kaszab 1952a: 574; Kwon and Choi 1986: Mozambique, CII.1994, A. Voehzkov leg. \ “G. juanensis
106; Hua Li 2002: 139; Iwan and Löbl 2008: 264. n. sp. Blair det.” in litt., 13 ex. (ZMHB), 1 ex. (JFCS).
Hopatrum japanum Motschulsky, 1860. – Gemminger and Harold Distribution. AFROTROPICAL REGION: Mozam-
1870: 1932. bique.
Opatrum expansicolle Lewis, 1894: 380.
Gonocephalum expansicolle (Lewis, 1894). – Gebien 1910b: 323;
Schuster 1928: 990; Reichardt 1936: 111; Kaszab 1952a: 678.
G. kandaharicum Kaszab, 1960
Type data. Unknown.
Gonocephalum kandaharicum Kaszab, 1960a: 148. – Kaszab 1968b:
Distribution. PALAEARCTIC REGION (Iwan and
20, 1970c: 152; Iwan and Löbl 2008: 264.
Löbl 2008): Asia: Japan, “East China”, “Korea”.
Remarks. S. China (Hua Li 2002: 139), South Korea
Type data . Holotype: allotype, 18 paratypes,
(Kwon and Choi 1986: 106).
“Afghanistan: Kandahar, 950 m, 13, 19, 22.II.1953, J.
Klapperich; Kandahar-Kuna, 950 m, 20, 28.I.1953, J.
Klapperich; Scham-Schir-Ror bei Kandahar, 950 m, 14.
G. javae Ferrer, 2010
I.1953, J. Klapperich; Koh-Bak bei Kandahar, 950 m, 13.
Gonocephalum javae Ferrer, 2010: 236 (replaced name). I.1953, J. Klapperich”.
Gonocephalum puberulum Kaszab, 1952a: 594. – Ferrer 2010: 236 Distribution. PALAEARCTIC REGION (Iwan and
(nom. preocc.: Gonocephalum puberulum Fauvel, 1867). Löbl 2008): Asia: Afghanistan.
Type data. Holotype: male (TMNH), “Madagaskar, Type data. Syntypes: 2 males, 2 females (BMNH),
Ankorokoka, Tulear dist. I.1956, C. Koch leg.”. SWA Hereroland, VIII–IX.1903, L. Schulze leg.
Paratypes: Madagaskar, Ankorokoka, Tulear dist. Distribution. AFROTROPICAL REGION: Namibia.
I.1956, C. Koch leg., 31 ex. (TMNH), 1 ex. (BMNH), 1 ex.
(JFCS); Tulear, id., 5 ex.; Androka Ampanihy, id., 1 ex.;
Marovato, Ambovombe distr. id., 1 ex. (BMNH); Tana- G. kondoi Masumoto, 1983
narive, Vadon leg., 1 ex. (MNHN); Madagascar “G. con-
Gonocephalum kondoi Masumoto, 1983: 173. – Iwan and Löbl 2008:
vexifrons Ferrer det.” in litt., 1 ex. (HNHM); Diego
264.
Suarez, Bontemps, ex coll. Perrier de la Bathie, 1906\
Gonocephalum n sp. J. Chatanay vidit.1915., 5 ex.
Type data. Unknown.
(MNHN).
Distribution. PALAEARCTIC REGION (Iwan and
Distribution. AFROTROPICAL REGION: Mada-
Löbl 2008): Asia: Japan (Ryukyus); Taiwan (Formosa).
gascar.
Type data . Holotype: male (MNHN), “Kikuyu Type data. Holotype (ZMHB), “Kosempo (leg. H.
Escarpement, 2.300 m., 1932-1933, Arambourg, Chap- Sauter)”.
puis et Jeannel, Miss. de l’Omo”. Distribution. PALAEARCTIC REGION (Iwan and
Distribution. AFROTROPICAL REGION: Kenya, Löbl 2008): Asia: Taiwan (Formosa).
Tanzania. Remarks. China: Fujian (Hua Li 2002: 139).
Gonocephalum Klapperichi Kaszab, 1952a: 550. – Kaszab 1954b: 248. Gonocephalum koreanum Kaszab, 1952a: 549. – Iwan and Löbl 2008:
Gonocephalum klapperichi Kaszab, 1952a. – Hua Li 2002: 139; Iwan 264.
and Löbl 2008: 264.
Type data. Holotype (HNHM), “Chemulpo”.
Type data. Syntypes: 3 ex. (HNHM). Distribution. PALAEARCTIC REGION (Iwan and
Material studied. “Japan, leg. Dönitz; Futschau; Löbl 2008): Asia: South Korea.
Kiautschau; Fukien, Shaovu 500 m (leg. J. Klapperich)”.
Other depositories. NHMB (2 ex.), MAKB (19 ex.).
Distribution. PALAEARCTIC REGION (Iwan and G. kuehnelti Kaszab, 1961
Löbl 2008): Asia: China: Fujian (Fukien), Shaanxi Gonocephalum Kühnelti Kaszab, 1961b: 351.
(Shensi); Japan. Gonocephalum kuehnelti Kaszab, 1961b. – Iwan and Löbl 2008: 264.
G. kolbei Gridelli, 1945 Gonocephalum kuluanum Kaszab, 1952a: 659. – Kaszab 1960a: 151,
1961a: 4, 1961b: 351,1961d: 236, 1968b: 20, 1970a: 424, 1970c: 153,
Gonocephalum kolbei Gridelli, 1945: 7. – Ferrer 1993a: 92, 1995a: 71. 1974a: 193; Kwieton 1981: 17; Iwan and Löbl 2008: 264.
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 277
Gonocephalum littorale Chatanay, 1914: 471. – Gridelli 1945: 4 (bona G. macleay Blackburn, 1907
species); Ferrrer 2000b: 103, 2004: 90.
Gonocephalum alticola var. littorale Chatanay, 1914: 471. – Gebien Gonocephalum Macleay Blackburn, 1907: 288. – Gebien 1910b: 323,
1939: 451. 1939: 449.
Type data. Lectotype (designated by Gridelli 1948: Type data. Syntypes (BMNH), “Toowoomba,
35): male (MNHN), “Kenya: Côte de l’Afrique orientale Queensland”.
Anglaise: Shimoni, Alluaud et Jeannel leg. “litorale Distribution. AUSTRALIAN AND PACIFIC RE-
Chat. typus! det. Gridelli 1948” “Gonocephalum litorale GIONS: Australia (Queensland).
Chat. spec. propries! Nec alticola var. det. Gridelli
1948”.
Distribution. AFROTROPICAL REGION: Kenia, G. macrophthalmum Kaszab, 1952
Mozambique.
Gonocephalum macrophthalmum Kaszab, 1952a: 582. – Kaszab
1980b: 171; Iwan and Löbl 2008: 264.
Gonocephalum lucidicolle Ardoin, 1965: 1321. – Ferrer 1993a: 84, Gonocephalum madurense Kaszab, 1952a: 625.
1995a: 33.
Type data. Syntypes: 10 ex. (HNHM).
Type data. Syntypes (MNHN), “Burkina Faso: Material studied. Madura; Shembaganur; Trichi-
Pabré, Haute Volta, R.P. Fernández leg.” nopoly, 1888 (leg. R. P. Castets).
Distribution. AFROTROPICAL REGION: Burkina Other depositories. BMNH (1 ex.), NHMB (1 ex.).
Faso, Guinea, Senegal, Sierra Leone. Distribution. ORIENTAL REGION: India.
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 279
G. martini Ferrer, 1995 Gonocephalum merensi estevezi Ferrer, 2000b: 71. – Iwan and Löbl
2008: 264.
Gonocephalum martini Ferrer, 1995a: 36. – Ferrer 2004: 86.
Type data. Holotype: male, “Gran Canaria, Punta
Type data. Holotype: male (BMNH), “Victoria West Arenas, 29.V.1989, M. A. Pena Estevez leg.”.
Cape Colony, 1948, P.D. Morris leg.”. Paratypes: id. 7.V.1989, (sub Launea arbores-
Paratypes: Victoria West Cape Colony, 1948, P.D. cens), 1 ex.; vegetation rastrea, 29.V.1989, coll. Peña
Morris leg., 1 female (BMNH); Basutoland, Dikolobeng Estévez, Servicio Agricola de Gran Canaria, 1 female
River 11 miles NE Quning, 16.III.1951 Brinck-Rudebeck (JFCS); Punta Arenas, 10.VII.1993, P. Oromí leg.,
leg., 2 females (ZMUL); Bon Accord Dam 10 miles 9 ex.
280 D. IWAN, J. FERRER and M. RAŚ
Distribution. PALAEARCTIC REGION (Iwan and Gonocephalum micantipenne Fairmaire, 1893. – Gebien 1910b: 324,
1939: 451; Chatanay 1913: 768; Ferrer 1993a: 80, 1995a: 23.
Löbl 2008): North Africa: Canary Islands (Gran
Canaria).
Type data. Syntypes: probably in MNHN.
Distribution. AFROTROPICAL REGION: Comores,
Madagascar, Mozambique, Seychelles, Somalia; Tan-
G. merensi merensi Uyttenboogaart, 1929 zania.
Gonocephalum merensi Uyttenboogaart, 1929: 348. – Gebien 1939:
446.
Gonocephalum merensi merensi Uyttenboogaart, 1929. – Ferrer G. michaelseni Gebien, 1920
1993a: 97, 2000b: 71 (stat. subsp.); Iwan and Löbl 2008: 264.
Gonocephalum Michaelseni Gebien, 1921 [1920]: 134. – Gebien 1939:
451.
Type data. Lectotype (designated by Ferrer 2000b:
Gonocephalum michaelseni Gebien, 1921. – Ferrer 1993a: 90, 1995a:
71): male (BMNH), “Gran Canaria, Bahia de Gando, 54.
X.1987, Rijksmuseet Leyden”.
Distribution. PALAEARCTIC REGION (Iwan and Type data. Type: “Namibie, Omaruru, Michelsen
Löbl 2008): North Africa: Canary Islands (Gran leg., coll. Gebien Hamburg”.
Canaria). Distribution. AFROTROPICAL REGION: Namibia.
Gonocephalum merkli Ferrer 2000b: 101. Opatrum minusculum Fairmaire, 1894: 20. – Champion 1895: 116.
Bradymerus minusculum Fairmaire, 1894: 20.
Gonocephalum minusculum (Fairmaire, 1894). – Gebien 1910b: 324,
Type data. Holotype: male (HNHM), “Tanzanie: Dar 1939: 448; Kaszab 1952a: 481.
es Salam. “Gonocephalum n. sp. det. Dr Kaszab 1982”. Bradymerus bidentulus Fairmaire, 1896: 23 – Gebien 1939: 448;
Distribution. AFROTROPICAL REGION: Tanzania Kaszab 1952a: 673.
(Dar es Salam).
Type data. Syntypes: probably in MNHN.
Distribution. ORIENTAL REGION: Burma, India.
PALAEARCTIC REGION (Iwan and Löbl 2008): Asia:
G. mesopotamicum Blair, 1923
India: Uttaranchal (Uttar Pradesh).
Gonocephalum mesopotamicum Blair, 1923: 123. – Schuster 1928:
989; Reichardt 1936: 108; Gebien 1939: 446; Kaszab 1965b: 669;
Iwan and Löbl 2008: 264. G. misellum Blackburn, 1907
Type data. Unknown. Gonocephalum misellum Blackburn, 1907: 288. – Gebien 1910b: 324,
Distribution. PALAEARCTIC REGION (Iwan and 1939: 449.
Gonocephalum modestum (Reiche, 1847). – Gemminger and Harold G. muelleri humerale Gridelli, 1945
1870: 1934 (= Opatrum prolixum Erichson, 1843); Reichardt
1936: 105 (= Opatrum prolixum Erichson, 1843); Gebien 1910b: Gonocephalum mülleri humeralis Gridelli, 1945: 13.
324 (= Opatrum prolixum Erichson, 1843), 1939: 445 (= Opa- Gonocephalum muelleri humerale Gridelli, 1945. – Ferrer 1993a:
trum prolixum Erichson, 1843); Ferrer 1993a: 78, 1995a: 13; Iwan 88, 1995a: 50, 1995b: 15, 1998a: 15.
and Löbl 2008: 264.
Gonocephalum inornatum Schuster, 1938: 61. – Gridelli 1948: 49;
Kaszab 1972: 379, 1982: 166; Ferrer 1993a: 78 (syn.), 1995a: 13. Type data. Syntype: «Typus»: 1 male (MCSN),
Gonocephalum obscurum inorantum Schuster 1938. – Español and “Somalie, Grabedarre, 1937, Fcca leg.”, designe lecto-
Viñolas 1983: 36. type.
Gonocephalum prolixum inornatum Schuster, 1938. – Gri-
Paratypes: Somalie, Grabedarre, 1937, Fcca leg.,
delli 1953: 61; Kaszab 1957: 61, 1972: 379, 1982a: 166; Grimm
1985: 18.
2 female (MCSN);
Opatrum sulcipenne Thomson, 1858: 84. Distribution. AFROTROPICAL REGION: Kenya,
Hopatrum sulcipenne Thomson, 1858. – Gemminger 1870: 1934. Somalia, Soudan.
Gonocephalum sulcipenne (Thomson, 1858). – Gebien 1910b: 324
(= Opatrum prolixum Erichson, 1843); Ferrer 1995a: 9 (= Opa-
trum prolixum Erichson, 1843).
G. muelleri muelleri Gridelli, 1945
Type data. Opatrum modestum Reiche, 1847: Gonocephalum muelleri Gridelli, 1945: 13. – Kaszab 1982b: 167; Iwan
2 syntypes “modestum Abyssinie/Reiche (NHRS). and Löbl 2008: 264.
Gonocephalum inornatum Schuster, 1938: Gonocephalum muelleri muelleri Gridelli, 1945. – Ferrer 1993a: 88,
1 male (ZSMB), Dire Daoua «Staud. inornat.»; Syntype: 1995a: 49; Lillig and Bremer 2002: 67.
1 male (ZSMB), Harrar (Abyss.) «Staud. inorn.».
Distribution. AFROTROPICAL REGION: Gabon, Type data. Paratype: Massaua, Erythrée, 5.XII.
Ethiopia, Somalia, Sudan. PALAEARCTIC REGION 1939, Cl. Müller leg., male (MCSN).
(Iwan and Löbl 2008): Asia: Iran; Saudi Arabia; Yemen Distribution. AFROTROPICAL REGION: Eritrea,
(incl. Socotra). Etiopia, Sudan, Somalia.
PALAEARCTIC REGION (Iwan and Löbl 2008):
Asia: Saudi Arabia; Yemen (incl. Socotra).
Type data. Paratypes: Soudan, Ed Damer Shendi, 1962b: 209, 1963: 207; Lindberg 1962: 47; Israelson et al. 1982: 128;
Oromí 1982: 272; Garcia 1991: 207; Ferrer 1993a: 76, 1995a: 5;
2–5.XI.1962, 1 male, Kassala, Haiya, 1–3.XII.1962,
Grimm 1995: 42; Iwan and Löbl 2008: 264.
1 male; Blue Nile, Wad Medani, 11–14.XI.1962, 2 ex.;
Kosti, 22.I.1963, R. Linnavuori leg., det. P. Ardoin,
Type data. Holotype: “Opatrum oblitum Woll. The
(ZMUH); Karthoum, 10.VII.1974, V. Seichrt leg., coll.
Canary Isl. T. V. Wollaston, B.M. 1864-80”.
J. Picka, 30 ex.; Wad Medani, 12.XII.1952, W.G. Buttik-
Distribution. PALAEARCTIC REGION (Iwan and
er leg., 28 ex. (TMNH).
Löbl 2008): North Africa: Canary Islands.
Distribution. AFROTROPICAL REGION: Sudan,
Uganda. PALAEARCTIC REGION (Iwan and Löbl
2008): Asia: Saudi Arabia.
G. oblongum (Fabricius, 1801)
1875: 231; Seidlitz 1894: 440 (syn. Opatrum prolixum Erichson, Type data. Syntypes: probably in MNHN.
1843); Reitter 1904: 145 (= Opatrum prolixum Erichson, 1843); Distribution. ORIENTAL REGION: India, Vietnam.
Gebien 1910b: 324 (= Opatrum prolixum Erichson, 1843), 1939:
445 (= Opatrum prolixum Erichson, 1843); Reichardt 1936: 105
PALAEARCTIC REGION (Iwan and Löbl 2008): Asia:
(= Opatrum prolixum Erichson, 1843); Grimm 1985: 17. China: Guandong (Kwantung), Hongkong, Yunnan;
Gonocephalum obscurum obscurum (Küster, 1849). – Español and South Korea; Taiwan (Formosa).
Viñolas 1983: 36 (stat. subsp.); Ferrer 1993a: 77, 1995a: 13; Lillig
and Bremer 2002: 66; Soldati 2010: 6; Iwan and Löbl 2008: 264.
Gonocephalum okinawanum Chujô, 1963: 149. – Kaszab 1964b: 40; G. parcesetosum Kaszab, 1952
Chûjô 1966: 7, 1978a: 63; Iwan and Löbl 2008: 264.
Gonocephalum parcesetosum Kaszab, 1952a: 516. – Schawaller
1997a: 10; Iwan and Löbl 2008: 264.
Type data. Holotype (FAKU), “Hentona, Okinava
Is., 28.IV.1958, K. Iha leg.”
Type data. Holotype (NHMB).
Paratypes: Hentona, Okinava Is., 28.IV.1958, K. Iha
Material studied. “Calcutta; Jorhat, Deseme, V.
leg., 9 ex., (Michitaka Chûjô)
1907”.
Distribution. PALAEARCTIC REGION (Iwan and
Other depository. HNHM (1 ex.).
Löbl 2008): Asia: Japan (Ryukyus).
Distribution. ORIENTAL REGION: India. PALAE-
ARCTIC REGION (Iwan and Löbl 2008): Asia: Nepal;
G. omoense Ardoin, 1967 “North India”.
Gonocephalum patrizii Gridelli, 1948: 23. Gonocephalum turchestanicum Gridelli, 1948: 9. – Kaszab 1960a:
Gonocephalum patrizii patrizii Gridelli, 1948. – Ferrer 1993a: 91, 147, 1961d: 236, 1968b: 20, 1970c: 150, 1974a: 193; Medvedev and
1995a: 55 (stat. subsp.), Lillig and Bremer 2002: 68. Nepesova 1985: 132; Hua Li 2002: 139.
Gonocephalum patruele turchestanicum Gridelli, 1948. – Ferrer
1993a: 88, 1995a: 46, 2010: 232; Iwan and Löbl 2008: 265.
Type data. Holotype: male (MCSN), “Somalie,
Belet Amin, 1935, Patrizi leg.”.
Distribution. AFROTROPICAL REGION: Ethiopia, Type data. Holotype: male (MCSN), “Turkestan,
Kenya, Somalia, Sudan. Issyk-Kue, Terski Tan, Vi.1902 coll. Hauser”.
Distribution. PALAEARCTIC REGION (Iwan and
Löbl 2008): Asia: Afghanistan; China: Xizang (Tibet),
G. patrizii lagosicum Ferrer, 1995 Kazakhstan; Turkmenistan.
Remark. PALEARCTIC REGION: Tajikistan, Uzbe-
Gonocephalum patrizii lagosicum Ferrer, 1995a: 55. kistan (Kaszab 1968b), Egypt: Sinai Peninsula (Lillig
and Pavlicek 2003).
Type data. Holotype: male (ZSMB),“Nigeria, Lagos
– sans autre indication Cl. Müller”.
Paratypes: Lagos – sans autre indication Cl. Müller, G. peguanum Kaszab, 1952
5 ex. (ZMS), 1 female id. (HNHM).
Distribution. AFROTROPICAL REGION: Nigeria. Gonocephalum peguanum Kaszab, 1952a: 639.
Gonocephalum famelicum (A. G. Olivier, 1811). – Mulsant and Rey Distribution. PALAEARCTIC REGION (Iwan and
1859: 70, 1859: 132; Baudi 1875: 229; Reitter 1904: 145; Reichardt
Löbl 2008): Asia: Afghanistan.
1936: 105; Gebien 1939: 444.
Opatrum persimile Lewis, 1894: 467. – Champion 1895: 117. Type data. Holotype: male (IRSN), “Ethiopie: prov.
Gonocephalum persimile (Lewis, 1894). – Gebien 1910b: 324, 1939: Goba, Bale, 1934-1935, C. de Mouleneare leg.”.
447; Schuster 1928: 990; Reichardt 1936: 113; Kaszab 1952a: 624, Paratypes: Ethiopie: prov. Goba, Bale, 1934-1935, C.
1968a: 8; Kwon and Choi 1986: 106; Hua Li 2002: 139; Iwan and Löbl
de Mouleneare leg., 5 ex. (IRSN), 1 ex. (JFCS);
2008: 265.
“Abyssinie” (Gonocephalum sp. aff. Alticola Chat. det.
Dr Kaszab), 1 ex. (HNHM); “Abyssinie” Gassner leg.
Type data. Unknown.
Coll. Staudinger (coll. Schuster), 12 ex. (ZSMB); Addis
Distribution. PALAEARCTIC REGION (Iwan and
Abeba, I.1939, E.v.Saalfeld leg., 1 ex. (ZSMB); Shoa
Löbl 2008): Asia: China: Heilongjiang (Heilungkiang),
prov. Valley of Chacha River, 55 km N Addis Abeba,
Nei Mongol (Inner Mongolia); Japan; Mongolia; North
14.VII.1973, R. Clarke leg., 10 ex. (MRAC); Valley of
Korea; Russia: Far East; South Korea.
Dima, 23.VII.1973, id., 1 ex. (MRAC); Akaki, 1XI.1900, 1
ex.; Warangambo, 18.II.1901, 1 ex.; Wollesch Wasam-
bara, 10–16.II.1901, 3 ex.; Fassasa Eksaua, 17.II.1901, 3
G. philippinense Kaszab, 1952
ex.; Nacmaoua, 17.II.1901, 1 ex. v. Erlanger leg., 1 ex.
Gonocephalum philippinense Kaszab, 1952a: 555. (BMHN); Prov. Wallega, Jabdo Kellega, 1.900 m.,
IV–V.1938, F.B. Nauhaus leg., 1 ex. (ZSMB); Harrar, M.
Type data. Holotype (NHMB), “Surigao, Mindanao de Rotschild Exp., 1 ex. (MNHN); Filoa Tchoba, id.
(leg. G. Böttcher)”. (MNHN); Schoa, Tscheratscha, O. Neumann leg., 1 ex.
Distribution. ORIENTAL REGION: Philippines. (ZMHB); Scioa, Argu-Ager, 1877, Antinori leg. “scioen-
sis milhi Gridelli det. 1957, 4 ex. (MCSN); Goggiam,
Ghiso-Abai, 15.IV.1939 (2.800 m), Cap Strobele leg.,
G. planicolle Kaszab, 1952 3 ex. (MCSN); Scioa, Mahal Vonz, Antinori leg., 1 ex.
(MCSN).
Gonocephalum planicolle Kaszab, 1952a: 468. Distribution. AFROTROPICAL REGION: Ethio-
pia.
Type data. Holotype (HNHM), “Bengalen, Raymahal”.
Distribution. ORIENTAL REGION: India.
G. pottsi Kulzer, 1957
Xizang (Tibet); Iran; Iraq; Kazakhstan; Mongolia; Abkhazia); Hungary; Italy (incl. Sardinia, Sicily, San
Oman; Pakistan; Saudi Arabia; Tajikistan; Turk- Marino); Russia: South European Territory; Slovakia;
menistan; Uzbekistan. Slovenia; Switzerland; Ukraine. Asia: China: Xizang
Remarks. Arab Emirates Union (Lillig and Bremer (Tibet); Kazakhstan; Turkey.
2002). Remarks. Turkmenistan (Reichardt 1936).
Gonocephalum pulverulentum Olivier, 1811: 159. Gonocephalum rastellii Ferrer, 1997: 212.
Type data. Unknown. Type data . Male, Sénégal, Gallel, St. Louis,
Distribution. PALAEARCTIC REGION: Asia centr, 25.V.1992, S. Rastelli le. Rec. G. Curletti (museo Civico
Caucas, Mediterranean Area, Mongolia. di Storia Naturale di Carmagnola).
Patratype: Sénégal, Gallel, St. Louis, 29.VII.1992
Rastelli. Coll. J. Ferrer.
G. pygmaeum (Steven, 1829) Distribution. AFROTROPICAL REGION: Senegal.
G. reticulatum Motschulsky, 1854 Kumaon, VI.1912; Pindar River, Garhwai, United Prov.,
VI.1937 (leg. J.C.M. Gardner); Nepal, Soondrijal;
Gonocephalum reticulatum Motschulsky, 1854: 47. – Reitter 1904:
Rotung, 1400 ft, 31.XII.1911 (leg. M. de Conrey), Yam-
141; Gebien 1910b: 324, 1939: 443; Reichardt 1936: 99; Kaszab
1952a: 485, 1968a: 8; Kwon and Choi 1986: 107; Medvedev 1990:
bung stream, 1100 ft, 17.I.1912 (leg. Kemp); Kumaun,
170; Medvedev and Lobanov 1990: 187; Hua Li 2002: 139; Iwan and Malwa Tal, 3600 ft, 7.V.1911 (leg. Kemp)”.
Löbl 2008: 265. Other depositories. BMNH (53 ex.), ZSMG (4 ex.).
Hopatrum reticulatum (Motschulsky, 1854). – Gemminger 1870: 1933. Distribution. PALAEARCTIC REGION (Iwan and
Gonocephalum mongolicum Reitter, 1889: 706. – Champion 1895:
Löbl 2008): Asia: India: Arunachal Pradesh Bhutan,
115; Reitter 1904: 141; Gebien 1910b: 324, 1939: 443; Schuster 1928:
989; Aoyama 1920; Medvedev 1973: 94.
Uttaranchal, Uttar Pradesh, Sikkim, Darjeeling Dis-
trict; Nepal. Oriental Region.
Type data. Unknown.
Distribution. PALAEARCTIC REGION (Iwan and
Löbl 2008): Asia: China: Beijing (Peking or Peiping), G. rostratum Bremer et Ferrer, 2002
Gansu (Kansu), Hebei (Hopeh), Heilongjiang (Heilung- Gonocephalum rostratum Bremer et Ferrer, 2002: 222.
kiang), Henan (Honana), Jiangsu (Kiangsu), Jilin
(Kirin), Ningxia (Ningsia), Nei Mongol (Inner Mongo- Type data. Holotype: male (Bremer coll. ZSMG),
lia), Qinghai (Tsinghai), Shaanxi (Shensi), Shandong “Burma, S Sagaing Division, Thandbody Paya env.
(Shantung), Shanxi (Shansi), Tianjin (Tsiensin); Mon- 19 km SE Moniwa, 19–20.VI.1997, J. Kalab leg.”
golia; North Korea; Russia: East Siberia. Paratypes: Burma, S Sagaing Division, Thandbody
Paya env. 19 km SE Moniwa, 19–20.VI.1997, J. Kalab
leg., 4 ex. (ZSMG), 4 ex. (MNHN), 2 ex. (HNHM), 4 ex.
G. rileyi Kaszab, 1952
(NHRS), 8 ex. (JFCS).
Gonocephalum Rileyi Kaszab, 1952a: 473. – Iwan and Löbl 2008: 265. Distribution. ORIENTAL REGION: Burma.
G. royi clarkei Ferrer, 1995 Dasus fuscus Herbst 1793. – Motschulsky 1849: 126.
Hopatrum fuscum Herbst, 1793. – Gemminger, 1870: 1931.
Gonocephalum royi clarkei Ferrer, 1995a: 44. – Ferrer 2004: 86. Gonocephalum fuscum (Herbst, 1793). – Brisout 1863: 45.
Opatrum fuscum Küster, 1849: 51.
Gonocephalum fuscum (Küster, 1849). – Seidlitz 1894: 444; Reitter
Type data. Holotype: male (HNHM), “Ethiopie, 1904: 146; Gebien 1910b: 326.
Gojeb River Valley, 80 km SW Jimma, 1600 m. Opatrum modestum Küster, 1849a: 53. – hom. nov. name preoccup.
24.VII.1971, R.O.S. Clarke leg. «Gonocephalum sp. aff nec Opatrum modestum Reiche, 1847: 369).
hispidocostatum Fairmaire, det. Dr Kaszab 1981»”. Hopatrum modestum Küster, 1849a. – Gemminger 1870: 1933.
Gonocephalum modestum (Küster, 1849a). – Seidlitz 1894: 444; Gebi-
Paratypes: Ethiopie, Gojeb River Valley, 80 km SW
en 1910b: 326.
Jimma, 1600 m. 24.VII.1971, R.O.S. Clarke leg. «Gono- Opatrum obductum Gebler, 1860: 38.
cephalum sp. aff hispidocostatum Fairmaire, det. Dr Hopatrum obductum Gebler, 1860. – Gemminger 1870: 1932.
Kaszab 1981», 1 female (HNHM); Abyssinie, Gibe River, Gonocephalum obductum (Gebler, 1860). – Reitter 1904: 146; Gebien
3.000 fts, 25.IV.1965, Ph. C. Zophiro leg., 1 male (BMNH). 1910b: 326.
Opatrum patruele Küster, 1849a: 51. – hom. nov. nom. preoccup. nec
Distribution. AFROTROPICAL REGION: Abys-
Opatrum patruele Erichson, 1843.
sinie, Ethiopia, Kenya. Hopatrum patruele Küster, 1849a. – Gemminger 1870: 1934 (=
Hopatrum vicinum Gemminger 1870).
Gonocephalum patruele (Küster, 1849a). – Brisout 1863: 45; Seidlitz
1894: 440; Kraatz 1873: 437.
G. rugatulum (Fairmaire, 1893)
Opatrum pulverulentum Olivier, 1811: 500.
Hopatrum rusticum var. pulverulentum Olivier, 1811. – Gem-
Bradymerus rugatulum Fairmaire, 1893: 23.
minger and Harold 1870: 1933.
Gonocephalum rugatulus (Fairmaire, 1893). – Gebien 1939: 448.
Gonocephalum pulverulentum Olivier, 1811. – Seidlitz 1894: 444;
Gonocephalum rugatulum (Fairmaire, 1893). – Kaszab 1952a: 476.
Gebien 1910b: 326.
Opatrum rusticum Brullé, 1832: 218. (nom. preoccup. nec Opatrum
Type data. Syntypes: probably in MNHN. rusticum Olivier, 1811: 498)
Distribution. ORIENTAL REGION: Burma, Laos, Opatrum setulosum Küster, 1849a: 47. – hom. nov. nom. preoccup.,
Thailand, Vietnam. nec Opatrum setulosum Faldermann, 1837.
Hopatrum setulosum Küster, 1849. – Gemminger and Harold 1870:
1934.
Gonocephalum setulosum (Küster, 1849). – Seidlitz 1894: 444; Reit-
G. rusticum (Olivier, 1811) ter 1904: 146; Gebien 1910b: 326.
Gonocephalum spoliatum Rey, 1892: 42 – Reitter 1904: 146.
Opatrum rusticum Olivier, 1811: 498. – Iwan and Löbl 2008: 736 Opatrum substriatum Besser, 1832: 18. – Küster 1849a: 49.
(nomen protectum). Hopatrum substriatum Besser, 1832. – Gemminger 1870: 1934.
Hopatrum rusticum Olivier, 1811. – Gemminger 1870: 1933. Gonocephalum substriatum (Besser, 1832). – Seidlitz 1894: 444;
Gonocephalum rusticum (Olivier, 1811). – Brulle 1832: 218; Lucas Gebien 1910b: 326.
1848: 334; Baudi 1875: 229; Mulsant 1854: 171; Jacqulin du Val Opatrum terrosum Küster, 1845: 28.
1861: 69; Brisout 1863: 45; Miedel 1880: 140; Rey 1892: 42; Seidlitz Hopatrum terrosum Küster, 1845. – Gemminger 1870: 1934.
1894: 444; Reitter 1904: 146, 1911: 335; Gebien 1910b: 325, 1939: Gonocephalum terrosum (Küster, 1845). – Seidlitz 1894: 444; Reitter
445; Muller 1920: 201; Schuster 1928: 989; Uyttenboogaart 1930: 1904: 146; Gebien 1910b: 326.
234, 1931: 11; Reichardt 1936: 106; Koch 1935: 93, 1937: 192; Hopatrum trichopterum Gemminger, 1870: 122. – Gemminger 1870:
Portevin 1934: 13; Kaszab 1940: 6, 1960a: 147, 1961a: 4, 1961c: 170, 1934.
1961d: 236, 1963a: 346,1967b: 551, 1968b: 20, 1970c: 150, 1973b: 56, Gonocephalum trichopterum (Gemminger, 1870). – Seidlitz 1894:
1974a: 193, 1979d: 268, 1982b: 166; Español 1947: 96, 1959: 111, 444; Gebien 1910b: 326.
1962a: 233, 1962b: 210; Lindberg 1962: 47; Skopin 1964:, 1968; Küh- Bolitophagus tricostatum Fischer von Waldheim, 1844: 128. – Schus-
nelt 1965: 36; Shalaby, El-Haidari and Derves 1966: 84; Medvedev ter 1934: 128.
1968, 1974, 1973: 94, 1990: 171; Canzoneri 1970: 76; Grimm 1981: 8, Gonocephalum tricostatum (Fischer von Waldheim, 1844). – Gebien
1985: 18, 1986b: 10, 1991: 28, 1995: 42; Kaluzhnaya 1982: 74; Oromí 1939: 446.
1982: 272; Medvedev and Nepesova 1985: 132; Canzoneri and Vien- Hopatrum vicinum Gemminger, 1870: 122. – Gemminger 1870:
na 1987: 20; Medvedev and Lobanov 1990: 187; Garcia 1991: 207; 1934.
Ferrer 1993a: 88, 1995a: 45; Chikatunov et al. 1997: 369; Ferrer and Gonocephalum vicinum (Gemminger, 1870). – Seidlitz 1894: 444;
Soldati 1999: 61; Hua Li 2002: 139; Leo and Fattorini 2002: 160; Lil- Gebien 1910b: 326.
lig and Bremer 2002: 67; Lillig and Pavlíček 2003: 50; Lo Cascio and Opatrum vestitum Küster, 1849a: 45.
Scupola 2004: 116; Iwan and Löbl 2008: 265; Soldati 2010: 6. Hopatrum vestitum Küster, 1849. – Gemminger 1870: 1934.
Opatrum errans Wollaston, 1854: 501. – Wollaston 1865: 412 (= Opa- Gonocephalum vestitum (Küster, 1849). – Gebien 1910b: 326.
trum fuscum Herbst, 1793). Opatrum villosum Steven, 1829: 97.
Hopatrum errans Wollaston, 1854. – Gemminger 1870: 1931. Hopatrum villosum Steven, 1829. – Gemminger 1870: 1933.
Gonocephalum errans (Wollaston, 1854). – Seidlitz 1894: 444; Gebien Gonocephalum villosum Steven, 1829: 97. – Seidlitz 1894: 444; Gebi-
1910b: 326. en 1910b: 326.
Opatrum famelicum Küster, 1849a: 48.
Gonocephalum famelicum (Küster, 1849). – Seidlitz 1894: 444; Gebi- Type data. Opatrum famelicum Küster, 1849.
en 1910b: 326.
Opatrum fuscum Herbst, 1793: 225. – Brulle 1838: 68; Küster 1849b:
“In Griechenland”.
51; Wollaston 1854: 500, 1864: 487, 1865: 412; Baudi 1876: 68; Iwan Opatrum errans Wollaston, 1854. “Habitat Mader-
and Löbl 2008: 736 (nomen oblitum). am, rariom exemplar in museo Loweano (a Dom.
290 D. IWAN, J. FERRER and M. RAŚ
Heinecken olim captum) conservatur, alterum in mon- G. schelpi Bremer et Ferrer, 1992
tibus prope Curral das Freiras vere novo A. D. 1848
Gonocephalum schelpi Bremer et Ferrer, 1992: 83.
egomet deprehensi, et duo specimina nuperrime com-
municavit Dom. Rousset.”
Opatrum fuscum Herbst, 1793. “Mus. Herbst” Type data. Holotype: male (Bremer coll. ZSMG),
Opatrum modestum Küster, 1849. “Im südlichen “Süd-Thailand, Umgebung Narativath, Mai 1983, ad
Frankreich. Beschreiben nach einem Dejeanischen lucem, Tawee Pongpaew leg.”.
Exemplar aus Sturm Sammlung”. Paratypes: Thailand, Phuket, in detritus of palm
Opatrum patruele Küster, 1849. “In Südfrank- vegetation at beach, 6.X.1990, Torbjörn Renquist leg.,
reich bei Montpellier. Nach dem Sturm’schen Original 1 ex (Bremer coll. ZSMG), 3 ex. (JFCS).
exemplar beschreiben”. Distribution. ORIENTAL REGION: Thailand.
Opatrum setulosum Küster, 1849. “Bei Carthage-
na in Spanien (Handschuch)”.
Opatrum substriatum Besser, 1832. “Ex Austria G. schneideri Reitter, 1898
superiori a D. Ullrich et nomine O. Linarioe Koll. e Dal-
Gonocephalum Schneideri Reitter, 1898: 14. – Reitter 1904: 142;
matia et ex insula Cephalonia adque Corfu a D. Par- Gebien 1910b: 326, 1939: 444; Schuster 1928: 989.
reiss teneo” Gonocephalum schneideri Reitter, 1898. – Reichardt 1936: 101;
Opatrum vestitum Küster, 1849. “Im südlichen Medvedev and Nepesova 1985: 131; Iwan and Löbl 2008: 265.
Europa”.
Distribution. AFROTROPICAL REGION: Algeria Type data. Unknown.
(Desert), Chad, Sudan. PALAEARCTIC REGION (Iwan Distribution. PALAEARCTIC REGION (Iwan and
and Löbl 2008): Europe: Albania; Armenia; Azerbaijan; Löbl 2008): Asia: Kazakhstan; Turkmenistan; Uzbe-
Azores; Croatia; France (incl. Corsica, Monaco); kistan.
Greece (incl. Crete); Italy (incl. Sardinia, Sicily, San
Marino); Macedonia; Malta; Spain (incl. Gibraltar);
Russia: South European Territory; Ukraine; North G. schusteri Kaszab, 1952
Africa: Canary Islands; Egypt; Libya; Morocco (incl.
Gonocephalum Schusteri Kaszab, 1952a: 507. – Kaszab 1980b: 171.
Western Sahara); Madeira Archipelago; Tunisia; Asia:
Afghanistan; China: Henan (Honana), Liaoning, Nei
Type data. Holotype (HNHM).
Mongol (Inner Mongolia), Shandong (Shantung),
Material studied. “Chien Hoa (leg. H. Fruhstorfer);
Xizang (Tibet); Egypt: Sinai Peninsula; Iran; Iraq;
Hanoi”.
Israel; Kazakhstan; Mongolia; Oman; Russia: East
Other depositories. NHMB (3 ex.), ZSMG (2 ex.).
Siberia; Saudi Arabia; Tajikistan; Turkey; Turk-
Distribution. ORIENTAL REGION: Vietnam.
menistan; Uzbekistan; .
Remarks. PALEARCTIC REGION: Jordan (Kath-
beh-Bader 1996), Syria (Seidlitz 1894), China: Xinjiang
(Sinkiang) (Hua Li 2002: 139). G. sculptithorax Gridelli, 1948
Type data. Holotype: male (MCSN), “Erythrèe, G. simplex hadroides (Wollaston, 1861)
Eritrea, Tessenei, IX, 1936, Remedelli leg. (“G.
Opatrum hadroides Wollaston, 1861: 215. – Wollaston 1877: 225.
andreinii n. sp. Gridelli det.”)”.
Hopatrum hadroides Wollaston, 1861. – Gemminger, 1870: 1932.
Paratypes: Asmara, 6.X.1968, K. V. v. Harde leg., 21 Gonocephalum hadroides (Wollaston, 1861). – Gebien 1910b: 326
ex. (SMS); Somalie. NW Tug Wajala, 1.500 m, (Haut (= Opatrum simplex Fabricius, 1801), 1939: 450 (= Opatrum
Plateau) 1–11.VIII.1980, H.J. Bremer leg., 1 ex.; (50.13, simplex Fabricius, 1801); Ardoin 1971, 1972.
N Erigavo, 4.950 fts. Hemming leg., 10 ex.; Thilbo Gonocephalum simplex hadroides (Wollaston, 1861). – Ferrer
1993a: 103 (stat. subsp.), 2000b: 91.
(T236) id., 14 ex. (TMNH); Afr.or.ital. Gimma, VII.1935,
G. Loro leg., 19 ex. (TMNH); Mogadiscio, Breuning leg.,
Type data. Lectotype (designated by Ferrer 2000b:
7 ex. (MRAC); Giballe, 10.XII.1982, S.B.S., (MZUF
91): male (BMNH), “St Helena, T. V. Wollaston, Brit.
8179); Hargeisa, 1.300m, 21–31.VII.1980, Verena Meyer
Mus. 1877.51, Opatrum hadroides Wollaston”.
leg., 9 ex. (ZSMB); Ethiopie: Maraca, Thery leg. (ex.
Paratypes: St Helena, T. V. Wollaston, Brit. Mus.
coll. Schuster), 2 ex.; id. samml. Müller, 1 ex.; Harar,
1877.51, Opatrum hadroides Wollaston, 2 ex. (BMNH).
Gassner (ex. coll. Schuster), 5 ex.; Soisa, Antiniri leg.,
Distribution. AFROTROPICAL REGION: St. Hele-
1 ex. (ZSMB); Harrar, VII.1922, G. Babuly; id. pr. de Dir
na Isl., Republic of South Africa.
Daoua, M. de Rotschild exp., 1 ex., Karsa, id., 15 ex.;
Bourka, id. Mieka, id., 2 ex.; Filoa Choba, id., 1 ex.;
Maro, 16.III.1909, A. Bonhoure leg., 1 ex.; Addis Abeba,
G. simplex jeanneli Chatanay, 1914
(S1057) don. Dr Halstead (no sorghum), 6 ex.
(ADAS\SLL); Djourjou Abaji, Didessa Goma, 2.000 fts, Gonocephalum Jeanneli Chatanay, 1914: 472.
10.II.1905, Ph. C. Zaphiro leg., 1 ex.; Ladjo, Arussi prov. Gonocephalum Jeannelli Chatanay, 1914. – Gebien 1939: 451.
10.II.1901, B. v. Erlanger leg., 16 ex.; Negele, 1.500 m, Gonocephalum simplex jeannelli Chatanay, 1914. – Ferrer 1993a:
101, 105 (stat. subsp.), 1994: 201, 1996a: 61, 2000b: 83.
5.VII.1976, A. Tsadaka leg. (chick pea), 1 ex. (BMNH);
Gambela, Ilubabor prov. 27.VIII.1972, R.O.S. Clarke
Type data. Lectotype (designated by Ferrer 2000b:
leg., 8 ex.; Nura, Arussi prov. 1.250 m, 29.IX.1984,
83): male (MNHN), “Ougande: Molo, Mau Escarpment,
Verena Meyer leg., 9 ex. (ZSMB), 2 ex. (JFCS); Bahar
Alluaud et Jeannel leg. XII.1911, 2.420 m. st 19 “G. jean-
Dar, 30 km SW Kl. Nil, 27.X.1968, H. Harde leg., 23 ex.
neli Chat, Gridelli det. Typus!”.
(SMNS), 4 ex. (JFCS); Koka Dam, Lago Koka (Addis
Paratype: Ougande: Molo, Mau Escarpment, Allu-
Abeba) 15–16.I.1989, L. Azzaroli et Tartaglia leg.,
aud et Jeannel leg. XII.1911, 2.420 m. st 19 G. jeanneli
26 ex. (MZUF), 7 ex. (JFCS); Djur Geb, Pachoda,
Chat, Gridelli det. Typus!, 1 male (MNHN).
Schweinfurth leg., 27 ex. (ZMHB); Holuyu Garobabe,
Distribution. AFROTROPICAL REGION: Burundi,
20.III.1901, v. Erlanger leg. Homologi Sangalo, 2 ex.;
Camerun, Central African Republic, Kenya, Tanzania,
Sonogoro Misa, id., 1 ex.; Umfusa, id., 8 ex.; Buba
Uganda, Urundi, Zaire.
Bua, id., 1 ex. (BMNH); Arsi-Asella, 2.400 m, II.1989,
Sven Person leg., 1 ex. (ZMUL); Arsi-Lole, 8.I.1989,
id., 78 ex. (ZMUL); Harrar, 14.V.1900, 7 ex.; Gan- G. simplex microphthalmum Ardoin, 1965
dali, Harraresa, 30.V.1900, 2 ex.; Kunsi, 6.VI.1900,
2 ex.; Wabbi, 14.VII.1900, 11 ex.; Dalota, 11.VIII.1900, Gonocephalum microphthalmum Ardoin, 1965: 1318. – Kaszab
(ZMHB). 1969a: 228; Girard 1975: 341.
Gonocephalum simplex microphthalmum Ardoin, 1965. – Ferrer
Distribution. AFROTROPICAL REGION: Djibouti,
1993a: 102 (stat. subsp.), 2000b: 87.
Ethiopie, Kenia, Sudan.
Type data. Paratypes: Sierra Leone, 1er relevé,
1.440 m, Eperons du Rénoncement, Miss. I.F.A.N.
24.V.1963, 1 male (ZSMB); Monts Loma, Piste de Ban-
G. simplex chatanayi Gebien, 1939
dahoro, Miss. I.F.A.N. id., 1 female (HNHM).
Gonocephalum simplex chatanayi Gebien, 1939. – Ferrer 1993a: Distribution. AFROTROPICAL REGION: Cameroon,
103 (stat. subsp.), 1998a: 359, 2000b: 92. Ivory Coast, Republic of Congo, Sierra Leone, Zaire.
Gonocephalum chatanayi Gebien, 1939: 451 (replaced name).
– Kaszab, 1971a: 227.
Gonocephalum persimile Chatanay, 1914: 467. – Gebien 1939: 451
G. simplex montisrolandi Ardoin, 1962
(nom. preocc.: Opatrum persimile Lewis, 1894: 381).
Distribution. AFROTROPICAL REGION: Gambia, Melut, Upper Nile, 17.2.1979, H.J. Bremer leg., 1 ex.
Guinea Bissau, Senegal. (ZSMB); Taufikia, 100 km S Melut, 25.XII.1913, F.
Konitzko leg., 29 ex. (BMNH); Karthoum, 10.VII.1974,
Seichrt leg., 18 ex. (coll. Picka, Prague); White Nile,
G. simplex segne (Thomson, 1858) Sobat River, Zaphiro leg., 1 ex.; Blue Nile, XII.1906, S.
Flower leg., 5 ex.; Singa Damazin, 15.XII.1962, Lin-
Opatrum segne Thomson, 1858: 85.
navuori, 1 ex.; Wad es Zahi, 10.V.1963, id.; An Nil Shen-
Hopatrum segne Thomson, 1858. – Gemminger 1870: 1933.
Gonocephalum segne (Thomson, 1858). – Gebien 1910b: 326 (= Opa-
di, 2.XI.1962, id., 4 ex.; Kordofan, Lake Neilah,
trum simplex Fabricius, 1801), 1939: 450 (= Opatrum simplex 11.II.1963, Iba Yambio, Equatoria prov, 16.IV.1963, 18
Fabricius, 1801). ex. (ZMUH); Aegypt. Soudan: Sennar, 19–28.II. Ebner
Gonocephalum simplex segne (Thomson, 1858). – Ferrer 1993a: 102 leg. “G. simplex var. inquinatum Schuster det.”
(stat. subsp.), 1996a: 61, 2000b: 83, 2001: 230.
(NHRS); Unter Adjuba, O. Neumann leg., 82 ex.
(ZMHB).
Type data. Lectotype (designated by Ferrer 2000b: Distribution. AFROTROPICAL REGION: Burkina
83): male (IRSN), “Coll RISCNB, Gabon, ex coll. Thom- Faso, Chad, Egypt, Ethiopia, Mali (Alfao), Senegal,
son; Op. segne Thoms. det. J. Thomson”. Sudan.
Distribution. AFROTROPICAL REGION: Came-
roon, Gabon, Ghana, Guinea, Guinea Bissau, Kenya,
Republic of Congo, Tanzania. G. simplex ubanguiense Ferrer, 2000
Type data. Holotype: male (MCSN), “Soudan, zone G. somalicum Gridelli, 1945
inondé, septembre 1946,, ex coll. I.F.A.N. («Diafarabé»)”.
Gonocephalum somalicum Gridelli, 1945: 19. – Ferrer, 1995: 69;
Paratype: Soudan, zone inondé, septembre 1946,
Schawaller 2004: 451; Iwan and Löbl 2008: 266.
1 male, ex coll. I.F.A.N. («Diafarabé»), 1 male (MNHN)
leg. Pitot.
Type data. Syntypes (MCSN): “Somalia Grabre-
Distribution. AFROTROPICAL REGION: Gambia,
darre,4.IV.1973, (Facca leg.)”
Guinea, Senegal, Sudan.
Distribution. AFROTROPICAL REGION: Somalia.
PALAEARCTIC REGION (Iwan and Löbl 2008): Asia:
Yemen (incl. Socotra), (Suqutra).
G. simulatrix (Fairmaire, 1891)
Gonocephalum soricinum soricinum (Reiche et Saulcy, 1857). Material studied. Trichinopoly; Dekkan (coll. Ind.
– Ferrer 1993a: 97 (stat. subsp.), 2000b: 74; Lillig and Pavlíček
Mus.); Madras (leg. Kirsch; leg. Capt. W. Patton);
2003: 50; Iwan and Löbl 2008: 266.
Pondichery (leg. Mouflet); Chaibasa (leg. Konietzko);
Calcutta; Ober-Assam (leg. Hartert); Coimbatore (leg.
Type data. Syntypes: probably in MNHN, “Bords de
Forst); Nedungadu, Tajore Distr., 28.II.1938 (leg. P. S.
la Mer Rouge”
Nathan); Mysore State, Shimoga (leg. P. S. Nathan);
Distribution. AFROTROPICAL REGION: Algeria
Madura Db., Arana Kurichi, 8.XII.1937 (leg. P. S.
(Desert), Benin, Chad, Erythrea, Ethiopia, Senegal,
Nathan); Salun, 2.VII.1938 (leg. P. S. Nathan); Kanara;
Sudan. ORIENTAL REGION: India. PALAEARCTIC RE-
Senji; Karachi (leg. T. R. Bell); Karachi, Sind (leg. Cum-
GION (Iwan and Löbl 2008): Asia: Afghanistan; Egypt:
ming); Bandra, 1905 (leg. Dr. Jayakar); Nilgiri Hills,
Sinai Peninsula; Iran; Israel; Jordan; Pakistan; Saudi
3500 ft, X.1910 (leg. H. L. Andrewes); Calcutta, V.1914,
Arabia; Syria; Yemen (incl. Socotra).
“at leight” (coll. Ind. Mus.); Calcutta, 12–13.II.1910 (leg.
Remarks. United Arab Emirates (Schawaller 2010:
Kemp); Malabar; Manaparai, S. India; Balugaon, Oris-
270).
sa, 26.I.1914, “at leight” (leg. N. Annandale); Bengal,
Purneah Distr., Katihar (leg. A. Paiva); Chota Nagpur,
Chakardhupur, 29.X.–1.X.1911 (leg. F. H. Gravely);
G. spangleri Kaszab, 1980 Bangalore (leg. Cameron); Bangalore, 3000 ft, 15.X.
1910 (leg. N. Annandale); Berhampur (leg. Atkinson);
Gonocephalum spangleri Kaszab, 1980: 140. – Kaszab 1979b: 63.
W. base of W. Ghats, Kulattupuzha, 19.XI.1908 (leg. N.
Annandale); Orissa, Puri, 24. bis 26.I.1911 (leg. N.
Type data. Holotype “C. Province, Kan District,
Annandale and F. H. Gravely); Orissa, Argul, 1913 (leg.
Hasalaka, Irrigation bungalow, 5 mi. NW of Mahiyan-
J. Taylor); Puri, Bhubanesvar, 4–6.XI.1912 (leg. N.
gana, 10.III–9.IV.1971, P, P, Spangler, Smithsonian
Annandale); Barkul, Chilka Lake, 25.I.1914, “at leight”
Institute, U.S. National Museum, Washington”.
(leg. N. Annandale); Ceylon (leg. Doflein); Süd-Ceylon
Paratyps “wie Holotypes, 4 ex.”.
(leg. H. Fruhstorfer); Trincomali (leg. W. Horn); Colom-
Distribution. ORIENTAL REGION: Sri Lanka.
bo (leg. W. Horn); Paradna (leg. W. Horn); Matale (leg.
W. Horn); Kandy (leg. W. Horn); Kandy, 16.IV.1908 (leg.
G. E. Bryant); Kandy, 22.V.1910 (coll. Ind. Mus.); Ban-
G. spinicolle (Fairmaire, 1896) darawella (leg. W. Horn); Puttalam (leg. W. Horn);
Weligama (leg. W. Horn); Amuradhapura (Low coun-
Bradymerus spinicolle Fairmaire, 1896: 22.
Gonocephalum spinicolle (Fairmaire, 1896). – Kaszab 1952a: 475,
try), 18–20.X.1911 (leg. C. F. C. Baker); Kanthalay, XI.
1979b: 62; Iwan and Löbl 2008: 266. 1915 (leg. C. F. C. Baker).
Other depositories. BMNH (97 ex.), ZMHB (22 ex.),
Type data. Unknown. NHMB (7 ex.), NMPC (205 ex.), SDEI (33 ex.), ZSMG
Distribution. ORIENTAL REGION: Sri Lanka. (16 ex.).
PALAEARCTIC REGION (Iwan and Löbl 2008): Asia: Distribution. ORIENTAL REGION: India, Sri Lan-
India: Himachal Pradesh. ka. PALAEARCTIC REGION (Iwan and Löbl 2008):
Asia: India: Kashmir, Uttaranchal (Uttar Pradesh);
Pakistan.
G. stevensi Kaszab, 1952
Type data. Holotype (NHMB), “Assam, Dejoo, Opatrum strangulatum Fairmaire, 1888: 128. – Champion 1895: 117.
North Lakhimpur, Base of Hills (leg. H. Stevens)”. Hopatrum strangulatum Fairmaire, 1888.
Gonocephalum strangulatum (Fairmaire, 1888). – Gebien 1910b:
Distribution. ORIENTAL REGION: India (Assam).
327, 1939: 446 (= Gonocephalum curvicolle Reitter, 1889);
Schuster 1928: 990; Reichardt 1936: 107 (= Gonocephalum cur-
vicolle Reitter, 1889); Kaszab 1952a: 684 (= Gonocephalum sex-
G. stoeckleini Kaszab, 1952 uale Marseul, 1876); Hua Li 2002: 139; Iwan and Löbl 2008: 266.
Gonocephalum Stöckleini Kaszab, 1952a: 662. – Kaszab 1961b: 351, Type data. Syntypes: probably in MNHN.
1979b: 66.
Gonocephalum stöckleini Kaszab, 1952. – Kaszab 1965a: 114.
Distribution. PALAEARCTIC REGION (Iwan and
Gonocephalum stoeckleini Kaszab, 1952. – Iwan and Löbl 2008: Löbl 2008): Asia: China: Beijing (Peking or Peiping),
266. Heilongjiang (Heilungkiang).
Remarks. Bangladesh; China: Hainan, Hubei,
Type data. Syntypes: 6 ex. (HNHM). Younnan; Japan; “Korea”; “Siberia” (Hua Li 2002: 139).
CATALOGUE OF THE WORLD GONOCEPHALUM SOLIER, 1834. PART 1 297
Opatrum strigosum Reiche, 1850: 369. Gonocephalum subsetosum erichsoni Ferrer, 2000b: 114.
Hopatrum strigosum Reiche, 1850. – Gemminger 1870: 1934; Baudi
1875: 230.
Type data. Holotype: male (TMNH), “Namibie: Kao-
Gonocephalum strigosum (Reiche, 1850). – Rey 1892: 42; Reitter 1904:
145; Gebien 1910b: 327, 1939: 445; Schuster 1928: 989, 1934: 181; koveld, Svartbois, Drift Kunene, VII. =1951, C. Koch leg.”.
Koch 1935: 93; Reichardt 1936: 105; Gridelli 1940 (1939): 242; Ardoin Paratypes: Namibie: Kaokoveld, Svartbois, Drift
1965: 74; Ferrer 1993a: 84, 1995a: 33; Iwan and Löbl 2008: 266. Kunene, VII. =1951, C. Koch leg., 3 ex. (TMNH);
Ongaria, VI.1954, Vernay Transvaal Museum Exp.,
Type data. Lectotype (desinated by Ardoine 1965: 1 ex. (TMNH); Angola: NE Moçamedes, R. Giraul 10 mil
74): female (MNHN), “Gonocephalum strigosum S, 27–29.II.1972, B.M. South leg., 3 ex. (BMNH).
(Reiche) lectotype female, P. Ardoin des. 1965”. Distribution. AFROTROPICAL REGION: Angola,
Distribution. AFROTROPIOCAL REGION: Daho- Namibia.
mey, Erithrea, Ethiopia. PALAEARCTIC REGION
(Iwan and Löbl 2008): North Africa: Algeria; Egypt;
Tunisia. Asia: Israel. G. subspinosum (Fairmaire, 1894)
Type data. Holotype: male (MCSN), “Afrique du Type data. Syntypes: probably in MNHN.
Sud: Transvaal, Zoutpansberg, 800 m, ex coll. Gebien”. Distribution. ORIENTAL REGION: Bangladesh,
Distribution. AFROTROPICAL REGION: Republic India, Indonesia (Sunda Isl.), Sri Lanka, Vietnam.
of South Africa, Zambezi. PALAEARCTIC REGION (Iwan and Löbl 2008): Asia:
Bhutan; China: Fujian (Fukien), Guandong (Kwan-
tung), Guizhou (Kweichow), Hubei (Hupeh), Hunan,
G. subrugulosum Reitter, 1887 Jiangsu (Kiangsu), Sichuan (Szechwan), Xizang
(Tibet), Yunnan; India: Arunachal Pradesh, Sikkim
Gonocephalum subrugulosum Reitter, 1887: 388. – Champion 1895:
115; Reitter 1904: 141; Gebien 1919b: 327, 1939: 443; Schuster 1928: (Darjeeling District); Nepal; Taiwan (Formosa).
989; Reichardt 1936: 98; Medvedev 1973: 94, 1990: 170; Medvedev Remarks . China: Guangxi (Kwangsi), Jiangxi
and Lobanov 1990: 187; Hua Li 2002: 139; Iwan and Löbl 2008: 266. (Kiangsi) (Hua Li 2002).
Gonocephalum coriaceum takara Nakane, 1963: 26. Gonocephalum tenuipes Kaszab, 1952a: 587. – Kaszab 1980b: 171;
Gonocephalum takara Nakane, 1963. – Kaszab 1946: 40 (bona spe- Schawaller 1997a: 12; Iwan and Löbl 2008: 266.
cies, stat. spec.), 1964b: 40; Chûjô 1966: 8; Iwan and Löbl 2008: 266.
Type data. Holotype (HNHM).
Type data. “Ryukyus”.. Material studied. Tenasserim; Burma, Mergui,
Distribution. PALAEARCTIC REGION (Iwan and I.1927 (leg. R. N. Parker); Tavoy (coll. In. Mus.).
Löbl 2008): Asia: Japan (Ryukyus). Other depository. BMNH (2 ex.).
Distribution. ORIENTAL REGION: Burma, India,
Vietnam. PALAEARCTIC REGION (Iwan and Löbl
G. tellinii Gridelli, 1945 2008): Asia: Nepal.
Type data. Paratypes: Ethiopie: Dire Daoua, Gonocephalum recticolle var. terminale Reichardt, 1936: 110, 210. –
Gebien 1939: 446.
Kovacs leg., 1 male. 1 female (HNHM); Harrar, Staud- Gonocephalum terminale Reichardt, 1936. – Kaszab 1952a: 577;
inger, inornatum Schuster and Gebien, 1 male (ZSMB); Kwon and Choi 1986: 106; Iwan and Löbl 2008: 266.
Distribution. AFROTROPICAL REGION: Djibuti,
Eritrea, Ethiopia, Somalia, Sudan. Type data. Syntypes: “Prov. Ussuriensis: flumen
PALAEARCTIC REGION (Iwan and Löbl 2008): Tjumen-ula ad limitem Koreae, 14.V.–23.VII.1913
Asia: Oman; Saudi Arabia; Yemen (incl. Socotra). (tsherskij!); Kamenj-Rybolov as lacum Chanka,
Remarks. United Arabian Emirates (Ferrer 1993a). 17–18.VII.1908 (idem!); Troitzkoe ad lacum Chanka,
5.IX.1914 (idem!); Okeanskaja prope Vladivostok,
6.VI.1914 (idem!); Korea: Paljman, 29.VI.1900 (P.
G. tenasserimicum Kaszab, 1952 Schmidt!)”.
Gonocephalum tenasserimicum Kaszab, 1952a: 602.
Distribution. PALAEARCTIC REGION (Iwan and
Löbl 2008): Asia: Japan; North Korea; Russia: Far
Type data. Syntypes: 4 ex. (HNHM). East; South Korea.
Material studied. Tranquebar; India or. (leg. Duld);
Tenasserim (leg. Helfer).
Other depositories. ZMHB (1 ex.), NHMB (1 ex.), G. thailandicum Kaszab et Chujô, 1966
NMPC (34 ex.), SDEI (3 ex.), ZSMG (1 ex.).
Gonocephalum thailandicum Kaszab et Chujô, 1966: 53.
Distribution. ORIENTAL REGION: India, Vietnam.
Type data. Syntypes: 3 ex. (BMNH), “Tibet”. Gonocephalum torridum (Champion, 1894: 359). – Blackburn 1896:
Distribution. PALAEARCTIC REGION (Iwan and 274; 1907: 287; Gebien 1910b: 327, 1939: 449.
G. tonkinense Kaszab, 1952 Gonocephalum tschilianum Kaszab, 1952a: 630. – Hua Li 2002: 139;
Iwan and Löbl 2008: 266.
Gonocephalum tonkinense Kaszab, 1952a: 532. – Kaszab 1970a: 423,
1980b: 171; Iwan and Löbl 2008: 266.
Type data. Syntypes: 2 ex. (HNHM), “Tschili”.
Distribution. PALAEARCTIC REGION (Iwan and
Type data. Syntypes: 5 ex. (NHMB).
Löbl 2008): Asia: China: Shanxi (Shansi).
Material studied. “Quang Yen (leg. R. Vitalis de Remarks. China: Fujian; Japan, “S. China” (Hua Li
Salvaza); Chiem Hoa (leg. H. Fruhstorfer)”.
2002: 139)
Other depositories. HNHM (1 ex.), ZSMG (3 ex.).
Distribution. ORIENTAL REGION: Vietnam. PALAE-
ARCTIC REGION (Iwan and Löbl 2008): Asia: Nepal. G. tuberculatum (Hope, 1831)
Type data. Gonocephalum quadrinodosum Gonocephalum vagum (Steven, 1829). – Gebien 1910b: 327, 1939: 448;
Reitter, 1904. Syntypes: 3 females, (Frey coll?), Kaszab 1952a: 662, 1979b: 66; Ferrer 1995a: 59; Hua Li 2002: 139;
Iwan and Löbl 2008: 266.
“China”. Hopatrum arenarium Fairmaire, 1894: 17. – Blair 1922: 269. hom.
Distribution. ORIENTAL REGION: Bangladesh, nov. nom. preoccup., nec Opatrum arenarium Fabricius, 1775.
Burma, Philippines, Vietnam. PALAEARCTIC REGION Gonocephalum arenarium (Fairmaire, 1894). – Kaszab 1952a: 684;
(Iwan and Löbl 2008): Asia: Afghanistan; China: Ferrer 1995a: 59 (syn.).
Opatrum planatum Walker, 1858: 284.
Hainan, Hongkong, Hubei (Hupeh), Sichuan (Szech-
Hopatrum planatum Walker, 1858. – Gemminger 1870: 1933.
wan), Yunnan; India: Kashmir; Himachal Pradesh, Gonocephalum planatum (Walker, 1858). – Gebien 1910b: 324, 1939:
Sikkim (Darjeeling District), Uttaranchal (Uttar 448 (= Opatrum moluccanum Blanchard, 1853); Blair 1922: 292;
Pradesh); Nepal; Pakistan; Taiwan (Formosa). Kaszab 1952a: 684; Ferrer 1995a: 59 (syn.).
Gonocephalum ingae Koch, 1955a: 527. Opatrum Walkeri Champion, 1894: 360. – Champion 1895: 117.
Gonocephalum veigai ingeai Koch, 1955. – Ferrer 1993a: 92 (stat. Gonocephalum Walkeri Champion, 1894: 360. – Gebien 1908a: 327,
subsp.), 1995a: 71. 1910b: 327.
Gonocephalum villiersi Gridelli, 1948: 17. – Ferrer 1993a: 84, 1995a: Gonocephalum zambesense Ferrer, 1995a: 67.
34; Lillig and Bremer 2002: 66.
Type data. Holotype: male (MNHN), “Mozambique,
Type data. Syntype: 1 male (MCSN), “Niger, Air, Zambeze, Env. De Chemba, Inhacoro, 1928 P. Lesne”.
Mts. Baguezans (1500–1600 m) VII.1947, Chopard leg.”. Paratypes: Mozambique, Zambeze, Env. De Chem-
Distribution. AFROTROPICAL REGION: Burkina ba, Inhacoro, 1928 P. Lesne, 1 male (JFCS); Inhacoro,
Faso, Niger, Senegal, Sudan. prés de Chemba, 1938, B. Thénot leg., 1 male (MNHN).
302 D. IWAN, J. FERRER and M. RAŚ
Distribution. AFROTROPICAL REGION: Mozam- 1947: 96, 1962a: 233, 1962b: 210; Kaszab 1963a: 346; Kühnelt 1965:
37; Oromí 1982: 272; Grimm 1986a: 76.
bique.
Opatrum virgatum Erichson, 1843: 249. – Reitter 1904: 144 (syn.
Opatropis hispida Brulle, 1838).
Hopatrum virgatum Erichson, 1843. – Gemminger 1870: 1934.
G. zoltani nom. nov.
Gonocephalum parallelum Kaszab, 1952a: 606. – Kaszab 1968b: 20, Type data. Opatrum affine Billberg, 1815. Lecto-
1970c: 149; Iwan and Löbl 2008: 264; hom. nov. nom. preoccup., nec type (designated by Ferrer 2000b: 123): female (NHRS),
Gonocephalum setulosum var. parallelum Normand, 1936: 363. “Ins. Caffraria; 13–14 O. affine Bilberg Nov. Act. Ups.
Gonocephalum parallelum Kaszab, 1952a: 606. – Kaszab 1968b: 20, VII.1815, f. 4.”
1970c: 149.
Paralectotype: Ins. Caffraria; 13–14 O. affine Bil-
berg Nov. Act. Ups. VII.1815, f. 4, female (NHRS).
Type data. Holotype (HNHM).
Opatrum virgatum Erichson, 1843. Syntypes: 3
Material studied. Assur (coll. F. Hauser), 1889 (leg.
ex. (ZMHB), “O. virgatum Erichson (48035) Angola,
Fischer); Indien; Quellgebiet des Indus, Vale Range,
Schönlein”.
Kalabagh (leg. Koken).
Distribution. AFROTROPICAL REGION: Angola,
Other depositories. BMNH (3 ex.), NHMA (3 ex.).
Cape Verde Is., Chad, Darfour, Gambia, Niger, Senegal,
Distribution. ORIENTAL REGION: India, Sri Lan-
Sudan. PALAEARCTIC REGION (Iwan and Löbl 2008):
ka. PALAEARCTIC REGION (Iwan and Löbl 2008):
Europe: Greece (incl. Crete); Spain (incl. Gibraltar).
Asia: Afghanistan; India: Kashmir; Iraq; Pakistan. North Africa: Canary Islands; Egypt; Morocco (incl.
Western Sahara); Madeira Archipelago. Asia: Israel.
Remarks. Yemen.
Subgenus Opatropis Reitter, 1904
Opatropis insulana insulana Ardoin, 1965: 80. – Ferrer 1993a: 107 Gonocephalum oculare Kaszab, 1952a: 622. – Ferrer 1993a: 107;
Gonocephalum (Opatropis) insulana insulana Ardoin, 1965. – Kaszab 1977: 257; Schawaller 1997a: 10; Iwan and Löbl 2008: 266.
Ferrer 2000b: 127 (stat. subsp.). Gonocephalum (Opatropis) oculare Kaszab, 1952a. – Ferrer 2000b:
128.