Unit 1
Unit 1
Unit 1
PALAEOANTHROPOLOGY*
Contents
1.0 Introduction
1.1 Definition
1.2 Aim of Paleoanthropology
1.3 Scope of Palaeoanthropology
1.4 Fossils and their Preservation
1.5 Process of Fossilization
1.5.1 Physico-Chemical Conditions for Fossilization
1.6 Significance of Fossils
1.7 Summary
1.8 References
1.9 Answers/Hints to Check Your Progress
Learning Objectives:
After reading this unit, you will be able to:
understand the definition, aim and subject matter of Palaeoanthropology;
comprehend the relationship of Palaeoanthropology with other
disciplines; and
know what are fossils, their process of preservation and fossilization and
significance of fossils.
1.0 INTRODUCTION
Paleoanthropology emerged as a science during the late nineteenth century. The
discovery of prehistoric artifacts in Pleistocene deposits soon led to the excavation
of fossilized human bones. The archaeologists and geologists who unearthed
these artifacts were primarily concerned with determining whether the human
fossils and the artifacts found by them actually dated from the Pleistocene, thus
offering evidence for the geological antiquity of humans. Prehistoric
archaeologists reconstructed the way of life of prehistoric peoples through the
artifacts found, while anthropologists examined the human fossils. They wanted
primarily to identify the races of prehistoric humans. It was within this context
that French anthropologists began to use the term “pale´o-anthropologie” to
refer to a new scientific discipline devoted to the study of prehistoric human
races and human paleontology.The field of paleoanthropology studies origin and
development of early humans as an interdisciplinary branch of anthropology
(Goodrum, 2014).
Fig. 1: Scientific disciplines that participate in the approach to reconstruct the process of
human evolution (After Henke and Rothe, 1994)
In the early beginning of paleoanthropology, the main question was quite simple:
is there a fossil record which proves the existence of our ancestors from ancient
10 times? The protagonists of paleoanthropology soon recognized the need for a
sophisticated empirical approach. The best basis for such a development existed Introducing
Palaeoanthropology
in France, where Pierre Marcelin Boule (1861-1942), a qualified geologist,
paleontologist, and archaeologist, unified in persona all necessary attributes to
establish the Discipline of Paleoanthropology. His classical processing of the
Neandertal skeletons from La Chapelle-aux-Saints (Boule 1911-1913) became
a landmark in the history of human palentology. He aimed to understand the
patterns of variation and the significance of anatomical differences. For this
reason, Boule, invented special instruments for qualification(s) and simple
statistical concepts to analyze the variation in human skeletons. Boule established
a palentology of humans, later on called paleoanthropology, as a scientific
discipline (Henke and Tattersall, 2007).
Check Your Progress
1) What is meant by Paleoanthropology?
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2) How does the field of Paleoanthropology reconstruct human past?
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1.2 AIM
Paleoanthropology is the study of human evolution and that of our closest living
relatives, the other primates. Humans, of course are primates and
paleoanthropologists recognize the importance of understanding primate evolution
as a necessary condition in understanding human evolution. This is the reason
primate evolution is most commonly considered a part of paleoanthropology as
opposed to the larger field of vertebrate paleontology. Paleoanthropology also
includes a variety of other fields that contributes to the study of various areas
like human evolution and variation. These include primate biology, systematics,
ecology, genetics and geology. And of course, since paleoanthropologists are
interested in the behavior of fossil humans, and since many of these humans left
material evidence of their behaviour in the fossil record, the analysis of this
record. Paleolithic archeology is also a major part of paleoanthropology. Because
the material evidence of the behavior of fossil humans is so ubiquitous, while it
is essentially non-existent in other animals, paleoanthropology is unique among
the historical sciences (Begun, 2013).
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Palaeoanthropology and
Primate Evolution 1.3 SCOPE OF PALEOANTHROPOLOGY
To adequately understand human bio-cultural evolution, we need a broad base
of information. Paleoanthropologists recover and interpret all the clues left by
early hominins. It is a diverse multidisciplinary pursuit that seeks to reconstruct
every possible bit of information concerning the dating, anatomy, behaviour,
and ecology of our hominin ancestors. In the past few decades, the study of
early hominins has marshaled the specialized skills of many different kinds of
scientists. This growing and exciting adventure includes, but is not limited to,
geologists, vertebrate paleontologists, archeologists, physical anthropologists and
paleoecologists.
Once identified, fossil beds likely to contain hominin finds become the focus for
further extensive surveying. For some sites, generally those postdating 2.6 mya
(the age of the oldest identified human artifacts), archaeologists take over in the
search for hominin material traces. We don’t necessarily have to find remains of
early hominins themselves to know that they consistently occupied a particular
area. Such material clues as artifacts inform us directly about early hominin
activities. Modifying rocks according to a consistent plan or simply carrying
them around from one place to another over fairly long distances (in a manner
not easily explained by natural means, like steam or glaciers is characteristic of
no other animal but hominins (Jurmain et al., 2011).
Fossils are the most important link between our past and present, in fact it’s
around fossil only that we can trace our evolutionary history. Fossils occur in
two forms: the actual remains of organisms, which are generally incomplete,
including three-dimensional molds of their external body, and the remains of an
animal’s activity, such as trackways. The remains of an animal’s activity are
called trace fossils (Cachel, 2015). Fossils are assessed by the techniques
of physical anthropology, comparative anatomy, and the theory of evolution.
Types of Preservation
The study of the formation and preservation of fossils is called taphonomy. There
are several different types of preservation and several different approaches to
classification. It is important to be able to understand the different ways that
fossils can be formed in order to know what they can look like. It is also worth
noting that a single fossil may fall into more than one of these categories.
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Palaeoanthropology and subjected to burial, compression, and differing chemical environments, the more
Primate Evolution
likely the original material is to be changed.
The older the fossil, the more likely it is to be in a state of altered preservation.
As sediments are buried deeper in the Earth and gradually lithify into rock, the
associated fossils also undergo alteration to a greater or lesser extent. Examples
of this type of preservation include permineralization, recrystallization,
replacement, formation of casts and or molds, and carbonization.
Permineralization, the most common form of alteration, is a process in which
porous organic structures, such as wood and bone, have their microscopic pore
spaces, left vacant by decay, filled by minerals precipitated from groundwater.
The original hard parts remain but are encased in extra material that fills in the
pores. The resulting fossil is heavier and denser than the original material. With
this type of fossilization, the fine details of microscopic structure are generally
preserved, occasionally even preserving details of cell structure. Petrified wood
is a fairly common example of permineralization.
Casts are formed when a mineral, sediment, or some other material fills in a
mold and hardens to form a copy of the original structure. True casts are relatively
rare in the fossil record, and internal molds (especially steinkerns) are often
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confused with them. A mold is a negative image of the original, while a cast is a Introducing
Palaeoanthropology
positive image, or duplicate of the original. Since molds and casts occur as a
result of the dissolution or destruction of the original material after the surrounding
matrix of sediment has hardened, only a limited amount of fine detail is usually
preserved.
There are, however, many other ways in which life-forms have left traces of
their existence. Sometimes insects were trapped in tree sap, which later became
hardened and chemically altered. Because inside the hardened amber there was
little or no oxygen, the insects have remained remarkably well preserved for
million of years, even with soft tissues and DNA still present. This fascinating
circumstance led author Michael Crichton to conjure the Jurassic Park novel
and motion pictures. Leaf imprints in hardened mud, or similar impressions of
small organisms, or even the traces of dinosaur feathers are fossils (Jurmain et
al., 2011).
Chemical conditions: Soon after death the soft parts of organism undergo
decomposition by the activity of bacteria. Similarly, organic matter associated
with the mineralized parts like proteinaceous material of shells, bones, tests etc.
often disappear, regarding these initially hard parts porous and brittle. Chemical
exchanges also occur between the decomposing organism and the sediment which
contains it. The organic matter engaged in this process infact results from
photosynthesis by phytoplankton; about a million tons of organic substances
having this origin are deposited annually in the oceans in present day conditions.
16 However, it has been known for some years that organic substance themselves
have been able to “fossilize” in many sediments and preservation of the substance Introducing
Palaeoanthropology
of the soft parts can occasionally occur. The constant refinement of methods of
investigation in chemical analysis makes it possible to reveal the preservation of
an ever increasing number of constituents of living matter. Palaeobiochemistry
has thus grown up, the results of which are of interest to various branches of
modern palentology. The absence of fossils in the Pre-Cambrian formations may
have resulted from the absence of proteins in the hard parts of the organisms
(Jain and Anantharaman, 1996).
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Introducing
1.7 SUMMARY Palaeoanthropology
1.8 REFERENCES
Begun, D. R. (2013). A Companion to Paleoanthropology. UK: Blackwell
Publishing Ltd.
3) Geologists do the initial survey to locate potential early hominin sites. Many
sophisticated techniques can contribute to this search, including aerial and
satellite photography. Paleontologists can also (through comparison with
known faunal sequences) give quick and dirty approximate ages of fossil
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Palaeoanthropology and sites in the field without having to wait for the expensive and time-consuming
Primate Evolution
chronometric analyses.
6) Fossil records are widely used in the study of the following: (a)
Chronostratigraphy (b) Biostratigraphy (c) Paleogeography (d) Palaeoclimate
(e) Palaeoecology and (f) Organic Evolution. For further details refer section
1.5.
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